Rough and smooth, hard and soft, moist and dry, hot and cold, itching, tickling, pricking, stinging, aching are skin sensations; but some of these are almost certainly compounds. The most successful way of isolating the elements out of these compounds is to explore the skin, point by point, with weak stimuli of different kinds. If a blunt metal point, or the point of a lead pencil, a few degrees cooler than the skin, is passed slowly over the skin, at most points no sensation except that of contact arises, but at certain points there is a clear sensation of cold. Within an area an inch square on the back of the hand, several of these cold spots can be found; and when the exploration is carefully made, and the cold spots marked, they will be found to give the same sensation every time. Substitute a metal point a few {198} degrees warmer than the skin, and a few spots will be found that give the sensation of warmth, these being the warmth spots. Use a sharp point, like that of a needle or of a sharp bristle, pressing it moderately against the skin, and you get at most points simply the sensation of contact, but at quite a number of points a small, sharp pain sensation arises. These are the pain spots. Finally, if the skin is explored with a hair of proper length and thickness, no sensation at all will be felt at most points, because the hair bends so readily when one end of it is pressed against the skin as not to exert sufficient force to arouse a sensation; but a number of points are found where a definite sensation of touch or contact is felt; these are the touch spots.

No other varieties of "spots" are found, and the four sensations of touch, warmth, cold and pain are believed to be the only elementary skin sensations. Itch, stinging and aching seem to be the same as pain. Tickle is touch, usually light touch or a succession of light touches. Smooth and rough are successions of touch sensations. Moist is usually a compound of smooth and cold. Hard and soft combine touch and the muscular sensation of resistance.

Hot and cold require more discussion. The elementary sensations are warmth and coolness, rather than hot and cold. Hot and cold are painful, and the fact is that strong temperature stimuli arouse the pain spots as well as the warmth or cold spots. Hot, accordingly, is a sensation compounded of warmth and pain, and cold a sensation composed of coolness and pain. More than this, when a cold spot is touched with a point heated well above the skin temperature (best to a little over 100 Fahrenheit), the curious fact is noted that the cold spot responds with its normal sensation of cold. This is called the "paradoxical cold sensation". From this fact it is probable that a hot object excites the cold sensation, along with those of warmth and {199} pain; so that the sensation of heat is a blend of the three. Another curious fact is that a very cold object produces a burning sensation indistinguishable from that of a hot object; so that the sensation of great cold, like that of heat, is probably a blend of the three elementary sensations of warmth, cold and pain.

Fig. 32.--Diagram of various sorts of sensory end-organ found in the skin.
A is a hair end-organ; the sensory axons can be seen coiling around the root of the hair; evidently a touch on the hair, outside, would squeeze the coiled axon and stimulate it. The hair is a bit of "accessory apparatus."
B is a touch corpuscle, consisting of a coiled axon-end surrounded by a little cone of other tissue.
C is an end-bulb, presumably belonging to the temperature sense. It has, again, a coiled axon-end surrounded by other tissue. The "coils" are really much more finely branched than the diagram shows.
D is a free-branched nerve end, consisting simply of a branched axon, with no accessory apparatus. It is the pain-sense organ.
E is a corpuscle of a type found in the subcutaneous tissue, as well as in more interior parts of the body. It contains an axon-end surrounded by a layered capsule.

The stimulus that arouses the touch sensation is a bending of the skin. That which arouses warmth or cold is of {200} course a temperature stimulus, but, strange as it may seem, the exact nature of the effective stimulus has not been agreed upon. Either it is a warming or cooling of the skin, or it is the existence of a higher or lower temperature in the skin than that to which the skin is at the moment "adapted". This matter will become clearer when we later discuss adaptation. The stimulus that arouses the pain sensation may be mechanical (as a needle prick), or thermal (heat or cold), or chemical (as the drop of acid), or electrical; but in any case it must be strong enough to injure or nearly to injure the skin. In other words, the pain sense organ is not highly sensitive, but requires a fairly strong stimulus; and thus it is fitted to give warning of stimuli that threaten injury.

Several kinds of sensory end-organ are found in the skin. There is the "spherical end-bulb", into which a sensory axon penetrates; it is believed to be the sense organ for cold. There is the rather similar "cylindrical end-bulb" believed to be the sense organ for warmth. There is the "touch corpuscle", found in the skin of the palms and soles, and consisting, like the end-bulbs, of a mass of accessory cells with a sensory axon ramifying inside it; this is an end-organ for the sense of touch. There is the hair end-organ, consisting of a sensory axon coiled about the root of the hair; this, also, is a touch receptor. Finally, there is the "free-branched nerve end", consisting simply of the branching of a sensory axon, with no accessory apparatus whatever; and this is the pain receptor. Perhaps the pain receptor requires no accessory apparatus because it does not need to be extremely sensitive.

Now since we find, in the skin, "spots" responsive to four quite different stimuli, giving four quite different sensations, and apparently provided with different types of end-organs, it has become customary to speak of four skin senses in place of the traditional "sense of touch". We {201} speak of the pain sense, the warmth sense, the cold sense, and the pressure sense, which last is the sense of touch proper.