COORDINATION

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The internal structure of nerve centers helps us see how coÖrdinated movement is produced. The question is, how {38} several muscles are made to work together harmoniously, and also how it is possible that a pin prick, directly affecting just a few sensory axons, causes a big movement of many muscles. Well, we find the sensory axon, as it enters the cord, sending off a number of side branches, each of which terminates in an end-brush in synaptic connection with the dendrites of a motor nerve cell.


Fig. 10.--CoÖrdination brought about by the branching of a sensory axon. (Figure text: cord, sensory neurone, motor neurone)

Thus the nerve current from a single sensory neurone is distributed to quite a number of motor neurones. Where there are central neurones in the arc, their branching axons aid in distributing the excitation; and so we get a big movement in response to a minute, though intense stimulus.

But the response is not simply big; it is definite, coordinated, representing team work on the part of the muscles as distinguished from indiscriminate mass action. That means selective distribution of the nerve current. The axons of the sensory and central neurones do not connect with any and every motor neurone indiscriminately, but link up with selected groups of motor neurones, and thus harness together teams that will work in definite ways, producing {39} flexion of a limb in the case of one such team, and extension in the case of another. Every reflex has its own team of motor neurones, harnessed together by its outfit of sensory and central neurones. The same motor neurone may however be harnessed into two or more such teams, as is seen from the fact that the same muscle may participate in different reflex movements; and for a similar reason we believe that the same sensory neurone may be utilized in more than one reflex arc.


Fig. 11.--CoÖrdination brought about by the branching of the axon of a central neurone. (Figure text: sensory, central, motor)

The most distinctive part of any reflex arc is likely to be its central neurones, which are believed to play the chief part in coÖrdination, and in determining the peculiarities of any given reflex, such as its speed and rhythm of action.

                                                                                                                                                                                                                                                                                                           

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