CHAPTER IV

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Vegetable Cells and their Structure—Stellate Tissues—Secondary Deposit—Ducts and Vessels—Wood-Cells—Stomata, or Mouths of Plants—The Camera Lucida, and Mode of Using—Spiral and Ringed Vessels—Hairs of Plants—Resins, Scents, and Oils—Bark Cells.

We will now suppose the young observer to have obtained a microscope and learned the use of its various parts, and will proceed to work with it. As with one or two exceptions, which are only given for the purpose of further illustrating some curious structure, the whole of the objects figured in this work can be obtained without any difficulty, the best plan will be for the reader to procure the plants, insects, etc., from which the objects are taken, and follow the book with the microscope at hand. It is by far the best mode of obtaining a systematic knowledge of the matter, as the quantity of objects which can be placed under a microscope is so vast that, without some guide, the tyro flounders hopelessly in the sea of unknown mysteries, and often becomes so bewildered that he gives up the study in despair of ever gaining any true knowledge of it. I would therefore recommend the reader to work out the subjects which are here mentioned, and then to launch out for himself on the voyage of discovery. I speak from experience, having myself known the difficulties under which a young and inexperienced observer has to labour in so wide a field, without any guide to help him to set about his work in a systematic manner.

The objects that can be most easily obtained are those of a vegetable nature, as even in London there is not a square, an old wall, a greenhouse, a florist’s window, or even a greengrocer’s shop, that will not afford an exhaustless supply of microscopic employment. Even the humble vegetables that make their daily appearance on the dinner-table are highly interesting; and in a crumb of potato, a morsel of greens, or a fragment of carrot, the enthusiastic observer will find occupation for many hours.

Following the best examples, we will commence at the beginning, and see how the vegetable structure is built up of tiny particles, technically called “cells.”

That the various portions of every vegetable should be referred to the simple cell is a matter of some surprise to one who has had no opportunity of examining the vegetable structure, and indeed it does seem more than remarkable that the tough, coarse bark, the hard wood, the soft pith, the green leaves, the delicate flowers, the almost invisible hairs, and the pulpy fruit, should all start from the same point, and owe their origin to the simple vegetable cell. This, however, is the case; and by means of a few objects chosen from different portions of the vegetable kingdom, we shall obtain some definite idea of this curious phenomenon.

I.

FIG.
1. Strawberry, cellular tissue 15. Wood-cells, Elder
2. Buttercup leaf, internal layer 16. Glandular markings and resin, “Cedar” pencil
3. Privet, Seed Coat, showing star-shaped cells 17. Do. Yew
4. Rush, Star-shaped cells 18. Scalariform tissue, Stalk of Fern
5. Mistletoe, cells with ringed fibre 19. Dotted Duct, Willow
6. Cells from interior of Lilac bud 20. Do. Stalk of Wheat
7. Bur-reed (Sparganium), square cells from leaf 21. Wood-cell, Chrysanthemum
8. Six-sided cells, from stem of Lily 22. Do. Lime-tree
9. Angular dotted cells, rind of Gourd 23. Dotted Duct, Carrot
10. Elongated ringed cells, anther of Narcissus 24. Cone-bearing wood, Deal
11. Irregular star-like tissue, pith of Bulrush 25. Cells, outer coat, Gourd
12. Six-sided cells, pith of elder 26. Ducts, Elm
13. Young cells from Wheat 27. Cellular tissue, Stalk of Chickweed
14. Do. rootlets of Wheat 28. Holly-berry, outer coat

I.

On PlateI. Fig.1, may be seen three cells of a somewhat globular form, taken from the common strawberry. Any one wishing to examine these cells for himself may readily do so by cutting a very thin slice from the fruit, putting it on a slide, covering it with a piece of thin glass (which may be cheaply bought at the optician’s, together with the glass slides on which the objects are laid), and placing it under a power of two hundred diameters. Should the slice be rather too thick, it may be placed in the live-box and well squeezed, when the cells will exhibit their forms very distinctly. In their primary form the cells seem to be spherical; but as in many cases they are pressed together, and in others are formed simply by the process of subdivision, the spherical form is not very often seen. The strawberry, being a soft and pulpy fruit, permits the cells to assume a tolerably regular form, and they consequently are more or less globular.

Where the cells are of nearly equal size, and are subjected to equal pressure in every direction, they force each other into twelve-sided figures, having the appearance under the microscope of flat six-sided forms. Fig.8, in the same Plate, taken from the stem of a lily, is a good example of this form of cell, and many others may be found in various familiar objects.

We must here pause for a moment to define a cell before we proceed further.

The cell is a close sac or bag formed of a substance called from its function “cellulose,” and containing certain semi-fluid contents as long as it retains its life. In the interior of the cell may generally be found a little dark spot, termed the “nÚcleus,” and which may be seen in Fig.1, to which we have already referred. The object of the nucleus is rather a bone of contention among the learned, but the best authorities on this subject consider it to be the vital centre of the cells, to and from which tends the circulation of the protoplasm, and which is intimately connected with the growth and reproduction of the cell. On looking a little more closely at the nucleus, we shall find it marked with several small light spots, which are termed “nuclÉoli.”

On the same Plate (Fig.2) is a pretty group of cells taken from the internal layer of the buttercup leaf, and chosen because they exhibit the series of tiny and brilliant green dots to which the colour of the leaf is due. The technical name for this substance is “chlorophyll,” or “leaf-green,” and it may always be found thus dotted in the leaves of different plants, the dots being very variable in size, number, and arrangement. A very fine object for the exhibition of this point is the leaf of AnÁcharis, the “Canadian timber-weed,” to be found in almost every brook and river. It also shows admirably the circulation of the protoplasm in the cell.

In the centre of the same Plate (Fig.12) is a group of cells from the pith of the elder-tree. This specimen is notable for the number of little “pits” which may be seen scattered across the walls of the cells, and which resemble holes when placed under the microscope. In order to test the truth of this appearance, the specimen was coloured blue by the action of iodine and dilute sulphuric acid, when it was found that the blue tint spread over the pits as well as the cell-walls, showing that the membrane is continuous over the pits.

Fig.7 exhibits another form of cell, taken from the SpargÁnium, or bur-reed. These cells are tolerably equal in size, and have assumed a square shape. They are obtained from the lower part of the leaf. The reader who has any knowledge of entomology will not fail to observe the similarity in form between the six-sided and square cells of plants and the hexagonal and square facets of the compound eyes of insects and crustaceans. In a future page these will be separately described.

Sometimes the cells take most singular and unexpected shapes, several examples of which will be briefly noticed.

In certain loosely made tissues, such as are found in the rushes and similar plants, the walls of the cells grow very irregularly, so that they push out a number of arms which meet each other in every direction, and assume the peculiar form which is termed “stellate,” or star-shaped tissue. Fig.3 shows a specimen of stellate tissue taken from the seed-coat of the privet, and rather deeply coloured, exhibiting clearly the beautiful manner in which the arms of the various stars meet each other. A smaller group of stellate cells taken from the stem of a large rush, and exemplifying the peculiarities of the structure, are seen in Fig.4.

The reader will at once see that this mode of formation leaves a vast number of interstices, and gives great strength with little expenditure of material. In water-plants, such as the reeds, this property is extremely valuable, as they must be greatly lighter than the water in which they live, and at the same time must be endowed with considerable strength in order to resist its pressure.

A less marked example of stellate tissue is given in Fig.11, where the cells are extremely irregular, in their form, and do not coalesce throughout. This specimen is taken from the pithy part of a bulrush. There are very many other plants from which the stellate cells may be obtained, among which the orange affords very good examples, in the so-called “white” that lies under the yellow rind, a section of which may be made with a very sharp razor, and placed in the field of the microscope.

Looking toward the bottom of the Plate, and referring to Fig.27, the reader will observe a series of nine elongated cells, placed end to end, and dotted profusely with chlorophyll. These are obtained from the stalk of the common chickweed. Another example of the elongated cell is seen in Fig.14, which is a magnified representation of the rootlets of wheat. Here the cells will be seen set end to end, and each containing its nucleus. On the left hand of the rootlet (Fig.13) is a group of cells taken from the lowest part of the stem of a wheat plant which had been watered with a solution of carmine, and had taken up a considerable amount of the colouring substance. Many experiments on this subject were made by the Rev. Lord S.G. Osborne, and may be seen at full length in the pages of the Microscopical Journal, the subject being too large to receive proper treatment in the very limited space which can here be given to it. It must be added that later researches have caused the results here described to be gravely disputed.

Fig.9 on the same Plate exhibits two notable peculiarities—the irregularity of the cells and the copiously pitted deposit with which they are covered. The irregularity of the cells is mostly produced by the way in which the multiplication takes place, namely, by division of the original cell into two or more new ones, so that each of these takes the shape which it assumed when a component part of the parent cell. In this case the cells are necessarily very irregular, and when they are compressed from all sides they form solid figures of many sides, which, when cut through, present a flat surface marked with a variety of irregular outlines. This specimen is taken from the rind of a gourd.

The “pitted” structure which is so well shown in this figure is caused by a layer of matter which is deposited in the cell and thickens its walls, and which is perforated with a number of very minute holes called “pits.” This substance is called “secondary deposit.” That these pits do not extend through the real cell-wall has already been shown in Fig.12.

This secondary deposit assumes various forms. In some cases it is deposited in rings round the cell, and is clearly placed there for the purpose of strengthening the general structure. Such an example may be found in the mistletoe (Fig.5), where the secondary deposit has formed itself into clear and bold rings that evidently give considerable strength to the delicate walls which they support. Fig.10 shows another good instance of similar structure; differing from the preceding specimen in being much longer and containing a greater number of rings. This object is taken from an anther of the narcissus. Among the many plants from which similar objects may be obtained, the yew is perhaps one of the most prolific, as ringed wood-cells are abundant in its formation, and probably aid greatly in giving to the wood the strength and elasticity which have long made it so valuable in the manufacture of bows.

Before taking leave of the cells and their remarkable forms, we will just notice one example which has been drawn in Fig.6. This is a congeries of cells, containing their nuclei, starting originally end to end, but swelling and dividing at the top. This is a very young group of cells (a young hair, in fact) from the inner part of a lilac bud, and is here introduced for the purpose of showing the great similarity of all vegetable cells in their earliest stages of existence.

Having now examined the principal forms of cells, we arrive at the “vessels,” a term which is applied to those long and delicate tubes which are formed of a number of cells set end to end, their walls of separation being absorbed.

In Fig.19 the reader will find a curious example of the “pitted vessel,” so called from the multitude of little markings which cover its walls, and are arranged in a spiral order. Like the pits and rings already mentioned, the dots are composed of secondary deposit in the interior of the tube, and vary very greatly in number, function, and dimensions. This example is taken from the wood of the willow, and is remarkable for the extreme closeness with which the dots are packed together.

Immediately on the right hand of the preceding figure may be seen another example of a dotted vessel (Fig.20), taken from a wheat stem. In this instance the cells are not nearly so long, but are wider than in the preceding example, and are marked in much the same way with a spiral series of dots. About the middle of the topmost cell is shown the short branch by which it communicates with the neighbouring vessel.

Fig.23 exhibits a vessel taken from the common carrot, in which the secondary deposit is placed in such a manner as to resemble a net of irregular meshes wrapped tightly round the vessel. For this reason it is termed a “netted vessel.” A very curious instance of these structures is given in Fig.26, at the bottom of the Plate, where are represented two small vessels from the wood of the elm. One of them—that on the left hand—is wholly marked with spiral deposit, the turns being complete; while, in the other instance, the spiral is comparatively imperfect, and the cell-walls are marked with pits. If the reader would like to examine these structures more attentively, he will find plenty of them in many familiar garden vegetables, such as the common radish, which is very prolific in these interesting portions of vegetable nature.

There is another remarkable form in which this secondary deposit is sometimes arranged that is well worthy of our notice. An example of this structure is given in Fig.18, taken from the stalk of the common fern or brake. It is also found in very great perfection in the vine. On inspecting the illustration, the reader will observe that the deposit is arranged in successive bars or steps, like those of a winding staircase. In allusion to the ladder-like appearance of this formation, it is called “scalariform” (Latin, scala, a ladder).

In the wood of the yew, to which allusion has already been made, there is a very peculiar structure, a series of pits found only in those trees that bear cones, and therefore termed the coniferous pitted structure. Fig.16 is a section of a common cedar pencil, the wood, however, not being that of the true cedar, but of a species of fragrant Juniper. This specimen shows the peculiar formation which has just been mentioned.

Any piece of deal or pine will exhibit the same peculiarities in a very marked manner, as is seen in Fig.24. A specimen may be readily obtained by making a very thin shaving with a sharp plane. In this example the deposit has taken a partially spiral form, and the numerous circular pits with which it is marked are only in single rows. In several other specimens of coniferous woods, such as the Araucaria, or Norfolk Island pine, there are two or three rows of pits.

A peculiarly elegant example of this spiral deposit may be seen in the wood of the common yew (Fig.17). If an exceedingly thin section of this wood be made, the very remarkable appearance will be shown which is exhibited in the illustration. The deposit has not only assumed the perfectly spiral form, but there are two complete spirals, arranged at some little distance from each other, and producing a very pretty effect when seen through a good lens.

The pointed, elongated shape of the wood-cells is very well shown in the common elder-tree (see Fig.15). In this instance the cells are without markings, but in general they are dotted like Fig.21, an example cut from the woody part of the chrysanthemum stalk. This affords a very good instance of the wood-cell, as its length is considerable, and both ends are perfect in shape. On the right hand of the figure is a drawing of the wood-cell found in the lime-tree (Fig.22), remarkable for the extremely delicate spiral markings with which it is adorned. In these wood-cells the secondary deposit is so plentiful that the original membranous character of the cell-walls is entirely lost, and they become elongated and nearly solid cases, having but a very small cavity in their centre. It is to this deposit that the hardness of wood is owing, and the reader will easily see the reason why the old wood is so much harder than the young and new shoots. In order to permit the passage of the fluids which maintain the life of the part, it is needful that the cell-wall be left thin and permeable in certain places, and this object is attained either by the “pits” described on page 43, or by the intervals between the spiral deposit.

At the right-hand bottom corner of PlateI. (Fig.28) may be seen a prettily marked object, which is of some interest. It is a slice stripped from the outer coat of the holly-berry, and is given for the purpose of illustrating the method by which plants are enabled to breathe the atmospheric air on which they depend as much as ourselves, though their respiration is slower. Among the mass of net-like cells may be seen three curious objects, bearing a rather close resemblance to split kidneys. These are the mouths, or “stÓmata,” as they are scientifically called.

In the centre of the mouths may be seen a dark spot, which is the aperture through which the air communicates with the passages between the cells in the interior of the structure. In the flowering plants their shape is generally rounded, though they sometimes take a squared form, and they regularly occur at the meeting of several surface cells. The two kidney-shaped cells which form the “mouth” are the “guard-cells,” so called from their function, since, by their change of form, they cause the mouth to open or shut, according to the needs of the plant. In young plants these guard-cells are very little below the surface of the leaf or skin, but in others they are sunk quite beneath the layer of cells forming the outer coat of the tissue. There are other cases where they are slightly elevated above the surface.

Stomata are found chiefly in the green portions of plants, and are most plentiful on the under side of leaves. It is, however, worthy of notice, that when an aquatic leaf floats on the water, the mouths are only to be found on the upper surface. These curious and interesting objects are to be seen in many structures where we should hardly think of looking for them; for instance, they may be found existing on the delicate skin which envelops the kernel of the common walnut. As might be expected, their dimensions vary with the character of the leaf on which they exist, being large upon the soft and pulpy leaves, and smaller upon those of a hard and leathery consistence. The reader will find ample amusement, and will gain great practical knowledge of the subject, by taking a plant, say a tuft of groundsel, and stripping off portions of the external skin or “epidermis” from the leaf or stem, etc., so as to note the different sizes and shapes of the stomata.

On the opposite bottom corner of PlateI. Fig.25, is an example of a stoma taken from the outer skin of a gourd, and here given for the purpose of showing the curious manner in which the cells are arranged about the mouth, no less than seven cells being placed round the single mouth, and the others arranged in a partially circular form around them.

Turning to PlateII., we find several other examples of stomata, the first of which (Fig.1) is obtained from the under surface of the buttercup leaf, by stripping off the external skin, or “epidermis,” as it is scientifically termed. The reader will here notice the slightly waved outlines of the cell-walls, together with the abundant spots of chlorophyll with which the leaf is coloured. In this example the stomata appear open. Their closure or expansion depends chiefly on the state of the weather; and, as a general rule, they are open by day and closed at night.

A remarkably pretty example of stomata and elongated cells is to be obtained from the leaf of the common iris, and may be prepared for the microscope by simply tearing off a strip of the epidermis from the under side of the leaf, laying it on a slide, putting a little water on it, and covering it with a piece of thin glass. (See PlateII. Fig.2.) There are a number of longitudinal bands running along the leaf where these cells and stomata appear. The latter are not placed at regular intervals, for it often happens that the whole field of the microscope will be filled with cells without a single stoma, whilst elsewhere a group of three or four may be seen clustered closely together.

Fig.3 on the same Plate exhibits a specimen of the beautifully waved cells, without mouths, which are found on the upper surface of the ivy leaf. These are difficult to arrange from the fresh leaf, but are easily shown by steeping the leaf in water for some time, and then tearing away the cuticle. The same process may be adopted with many leaves and cuticles, and in some cases the immersion must be continued for many days, and the process of decomposition aided by a very little nitric acid in the water, or by boiling.

On the same Plate are three examples of spiral and ringed vessels, types of an endless variety of these beautiful and interesting structures. Fig.4 is a specimen of a spiral vessel taken from the lily, and is a beautiful example of a double spire. The deposit which forms this spiral is very strong, and it is to the vast number of these vessels that the stalk owes its well-known elasticity. In many cases the spiral vessels are sufficiently strong to be visible to the naked eye, and to bear uncoiling. For example, if a leaf-stalk of geranium be broken across, and the two fragments gently drawn asunder, a great number of threads, drawn from the spiral vessels, will be seen connecting the broken ends. In this case the delicate membranous walls of the vessel are torn apart, and the stronger fibre which is coiled spirally within it unrolls itself in proportion to the force employed. In many cases these fibres are so strong that they will sustain the weight of an inch or so of the stalk.

In Fig.5 is seen a still more bold and complex form of this curious structure; being a coil of five threads, laid closely against each other, and forming, while remaining in their natural position, an almost continuous tube. This specimen is taken from the root of the water lily, and requires some little care to exhibit its structure properly.

Every student of nature must be greatly struck with the analogies between different portions of the visible creation. These spiral structures which we have just examined are almost identical in appearance, and to some extent in their function, with the threads that are coiled within the breathing tubes of insects. This is in both cases twofold, namely, to give support and elasticity to a delicate membrane, and to preserve the tube in its proper form, despite the bending to which it may be subjected. When we come to the anatomy of the insect in a future page we shall see this structure further exemplified.

In some cases the deposit, instead of forming a spiral coil, is arranged in a series of rings, and the vessel is then termed “annulated.” A very good example of this formation is given in Fig.6, which is a sketch of such a vessel, taken from a stalk of the common rhubarb. To see these ringed vessels properly, the simplest plan is to boil the rhubarb until it is quite soft, then to break down the pulpy mass until it is flattened, to take some of the most promising portions with the forceps, lay them on the slide and press them down with a thin glass cover. They will not be found scattered at random through the fibres, which elsewhere present only a congeries of elongated cells, but are seen grouped together in bundles, and with a little trouble may be well isolated, and the pulpy mass worked away so as to show them in their full beauty. As may be seen in the illustration, the number of the rings and their arrangement is extremely variable. A better, but somewhat more troublesome, plan is to cut longitudinal sections of the stem, as described in our concluding chapter, when not only the various forms of cells and vessels, but their relations to each other, will be well shown. The numerous crystals of oxalate of lime, which make rhubarb so injurious a food for certain persons, will also be well seen. These crystals are called “raphides,” and are to be found in very many plants in different forms.

II.

FIG.
1. Cuticle, Buttercup leaf 24. Do. Pine cone
2. Do. Iris 25. Vitta, Caraway Seed
3. Do. Ivy leaf 26. Cork
4. Spiral vessel, Lily 27. Hair, Flower of Garden Verbena
5. Do. root, (rhizome) Water Lily 28. Do. fruit of Plane
6. Ringed vessel, Rhubarb 29. Do. do.
7. Chaff, after burning 30. Do. do.
8. Bifid hair, Arabis 31. Do. Lobelia
9. Hair, Marvel of Peru 32. Do. Cabbage
10. End of hair, leaf of Hollyhock 33. Do. Dead-nettle flower
11. Hair, Sowthistle leaf 34. Do. Garden Verbena flower
12. Do. Tobacco 35. Fruit-hair, Dandelion
13. Do. Southernwood 36. Hair, Thistle leaf
14. Group of hairs, Hollyhock leaf 37. Do. Cactus
15. Hair, Yellow Snapdragon 38. Do. do.
16. Do. Moneywort 39. Do. Virginian Spider-wort
17. Hair, Geum 40. Do. Lavender
18. Do. Flower of Heartsease 41. Section, Lavender leaf, Hairs
19. Do. Dockleaf and perfume-gland
20. Do. Throat of Pansy 42. Section, Orange Peel
21. Do. Dead-nettle Flower 43. Sting of Nettle
22. Do. Groundsel 44. Hair, Marigold flower
23. Cell, Beech-nut 45. Do. Ivy

II.

The hairs of plants form very interesting objects, and are instructive to the student, as they afford valuable indications of the mode in which plants grow. They are all appendages of and arise from the skin or epidermis; and although their simplest form is that of a projecting and elongated cell, the variety of shapes which are assumed by these organs is inexhaustible. On PlateII. are examples of some of the more striking forms, which will be briefly described.

The simple hair is well shown in Figs.18, 19, and 32, the first being from the flower of the heartsease, the second from a dock-leaf, and the third from a cabbage. In Fig.18 the hair is seen to be but a single projecting cell, consisting only of a wall and the contents. In Fig.19 the hair has become more decided in shape, having assumed a somewhat dome-like form; and in Fig.32 it has become considerably elongated, and may at once be recognised as a true hair.

In Fig.8 is a curious example of a hair taken from the white Arabis, one of the cruciferous flowers, which is remarkable for the manner in which it divides into two branches, each spreading in opposite directions. Another example of a forked hair is seen in Fig.13, but in this instance the hair is composed of a chain of cells, the three lower forming the stem of the hair, and the two upper being lengthened into the lateral branches. This hair is taken from the common southernwood.

In most cases of long hairs, the peculiar elongation is formed by a chain of cells, varying greatly in length and development. Several examples of these hairs will be seen on the same Plate.

Fig.9 is a beaded hair from the Marvel of Peru, which is composed of a number of separate cells placed end to end, and connected by slender threads in a manner that strongly reminds the observer of a chain of beads strung loosely together, so as to show the thread by which they are connected with each other. Another good example is seen at Fig.11, in a hair taken from the leaf of the sowthistle. In this case the beads are strung closely together, and when placed under a rather high power of the microscope have a beautifully white and pearly aspect. The leaf must be dry and quite fresh, and the hairs seen against the green of the leaf. Fig.39 represents another beaded hair taken from the Virginian Spiderwort, or Tradescantia. This hair is found upon the stamens, and is remarkable for the beautifully beaded outline, the fine colouring, and the spiral markings with which each cell is adorned.

A still further modification of these many-celled hairs is found in several plants, where the hairs are formed by a row of ordinarily shaped cells, with the exception of the topmost cell, which is suddenly elongated into a whip-like form. Fig.22 represents a hair of this kind, taken from the common groundsel; and Fig.36 is a still more curious instance, found upon the leaf of the thistle. The reader may have noticed the peculiar white “fluffy” appearance of the thistle leaf when it is wet after a shower of rain. This appearance is produced by the long lash-like ends of the hairs, which are bent down by the weight of the moisture, and lie almost at right angles with the thicker portions of the hair.

An interesting form of hair is seen in the “sting” of the common nettle. This may readily be examined by holding a leaf edgewise in the stage forceps, and laying it under the field of the microscope. In order to get the proper focus throughout the hair, the finger should be kept upon the screw movement, and the hair brought gradually into focus from its top to its base. The general structure of this hair is not unlike that which characterises the fang of a venomous serpent. The acrid fluid which causes the pain is situated in the enlarged base of the hair, and is forced through the long straight tubular extremity by means of the pressure exerted when the sting enters the skin. At the very extremity of the perfect sting is a slight bulb-like swelling, which serves to confine the acrid juice, and which is broken off on the least pressure. The sting is seen in Fig.43.

The extremities of many hairs present very curious forms, some being long and slender, as in the examples already mentioned, while others are tipped with knobs, bulbs, clubs, or rosettes in endless variety.

Fig.12 is a hair of the tobacco leaf, exhibiting the two-celled gland at the tip, containing the peculiar principle of the plant, known by the name of “nicotine.” The reader will see how easy it is to detect adulteration of tobacco by means of the microscope. The leaves most generally used for this purpose are the dock and the cabbage, so that if a very little portion of leaf be examined the character of the hairs will at once inform the observer whether he is looking at the real article or its substitute.

Fig.15 is a hair from the flower of the common yellow snapdragon, which is remarkable for the peculiar shape of the enlarged extremity, and for the spiral markings with which it is decorated. Fig.16 is a curious little knobbed hair found upon the moneywort, and Fig.17 is an example of a double-knobbed hair taken from the Geum. Fig.34 affords a very curious instance of a glandular hair, the stem being built up of cells disposed in a very peculiar fashion, and the extremity being developed into a beautiful rosette-shaped head. This hair came from the Garden Verbena.

Curiously branched hairs are not at all uncommon, and some very good and easily obtained examples are given on PlateII.

Fig.28 is one of the multitude of branched hairs that surround the well-known fruit of the plane-tree, the branches being formed by some of the cells pointing outward. These hairs do not assume precisely the same shape; for Fig.29 exhibits another hair from the same locality, on which the spikes are differently arranged, and Fig.30 is a sketch of another such hair, where the branches have become so numerous and so well developed that they are quite as conspicuous as the parent stem.

One of the most curious and interesting forms of hair is that which is found upon the lavender leaf, and which gives it the peculiar bloom-like appearance on the surface.

This hair is represented in Figs.40 and 41. On Fig.40 the hair is shown as it appears when looking directly upon the leaf, and in Fig.41 a section of the leaf is given, showing the mode in which the hairs grow into an upright stem, and then throw out horizontal branches in every direction. Between the two upright hairs, and sheltered under their branches, may be seen a glandular appendage not unlike that which is shown in Fig.16. This is the reservoir containing the perfume, and it is evidently placed under the spreading branches for the benefit of their shelter. On looking upon the leaf by reflected light the hairs are beautifully shown, extending their arms on all sides; and the globular perfume cells may be seen scattered plentifully about, gleaming like pearls through the hair-branches under which they repose. They will be found more numerous on the under side of the leaf.

This object will serve to answer a question which the reader has probably put to himself ere this, namely, Where are the fragrant resins, scents, and oils stored? On PlateI. Fig.16, will be seen the reply to the first question; Fig.41 of the present Plate has answered the second question, and Fig.42 will answer the third. This figure represents a section of the rind of an orange, the flattened cells above constituting the delicate yellow skin, and the great spherical object in the centre being the reservoir in which the fragrant essential oil is stored. The covering is so delicate that it is easily broken, so that even by handling an orange some of the scent is sure to come off on the hands, and when the peel is stripped off and bent double, the reservoirs burst in myriads, and fling their contents to a wonderful distance. This may be easily seen by squeezing a piece of orange peel opposite a lighted candle, and noting the distance over which the oil will pass before reaching the flame, and bursting into little flashes of light. Other examples are given on the same plate.

Returning to the barbed hairs, we may see in Fig.35 a highly magnified view of the “pappus” hair of a dandelion, i.e. the hairs which fringe the arms of the parachute-like appendage which is attached to the seed. The whole apparatus will be seen more fully on PlateIII. Figs.44, 45, 46. This hair is composed of a double layer of elongated cells lying closely against each other, and having the ends of each cell jutting out from the original line. A simpler form of a double-celled, or more properly a “duplex” hair, will be seen in Fig.44. This is one of the hairs from the flower of the marigold and has none of the projecting ends to the cells.

In some instances the cell-walls of the hairs become greatly hardened by secondary deposit, and the hairs are then known as spines. Two examples of these are seen in Figs.37 and 38, the former being picked from the Indian fig-cactus, and well known to those persons who have been foolish enough to handle the fig roughly before feeling it. The wounds which these spines will inflict are said to be very painful, and have been compared to those produced by the sting of the wasp. The latter hair is taken from the Opuntia. These spines must not be confounded with thorns; which latter are modified branches.

Fig.10 represents the extreme tip of a hair from the hollyhock leaf, subjected to a lens of very high power.

Many hairs assume a star-like appearance, an aspect which may be produced in different ways. Sometimes a number of simple hairs start from the same base, and by radiating in different directions produce the stellate effect. An example of this kind of hair may be seen in Fig.14, which is a group of hairs from the hollyhock leaf. There is another mode of producing the star-shape which may be seen in Fig.45, a hair taken from the leaf of the ivy. Very fine examples may also be found upon the leaf of Deutzia scabra.

Hairs are often covered with curious little branches or protuberances, and present many other peculiarities of form which throw a considerable light upon certain problems in scientific microscopy.

Fig.33 represents a hair of two cells taken from the flower of the well-known dead-nettle, which is remarkable for the number of knobs scattered over its surface. A similar mode of marking is seen in Fig.31, a club-shaped hair covered with external projections, found in the flower of the Lobelia. In order to exhibit these markings well, a power of two hundred diameters is needed. Fig.21 shows this dotting in another hair from the dead-nettle, where the cell is drawn out to a great length, but is still covered with these markings.

Fig.20 is an example of a very curious hair taken from the throat of the pansy. This hair may readily be obtained by pulling out one of the petals, when the hairs will be seen at its base. Under the microscope it has a particularly beautiful appearance, looking just like a glass walking-stick covered with knobs, not unlike those huge, knobby club-like sticks in which some farmers delight, where the projections have been formed by the pressure of a honeysuckle or other climbing plant.

A hair of a similar character, but even more curious, is found in the same part of the flower of the Garden Verbena (see Fig.27), and is not only beautifully translucent, but is coloured according to the tint of the flower from which it is taken. Its whole length is covered with large projections, the joints much resembling the antennÆ of certain insects; and each projection is profusely spotted with little dots, formed by elevation of the outer skin or cuticle. These are of some value in determining the structure of certain appearances upon petals and other portions of the flowers, and may be compared with Figs.33 to 35 on PlateIII.

Fig.26 offers an example of the square cells which usually form the bark of trees. This is a transverse section of cork, and perfectly exhibits the form of bark cells. The reader is very strongly advised to cut a delicate section of the bark of various trees, a matter very easily accomplished with the aid of a sharp razor and a steady hand.

Fig.24 is a transverse section through one of the scales of a pine-cone, and is here given for the purpose of showing the numerous resin-filled cells which it displays. This may be compared with Fig.16 of PlateI. Fig.25 is a part of one of the “vittÆ,” or oil reservoirs, from the fruit of the caraway, showing the cells containing the globules of caraway oil. This is rather a curious object, because the specimen from which it was taken was boiled in nitric acid, and yet retained some of the oil globules. Immediately above it may be seen (Fig.23) a transverse section of the beechnut, showing a cell with its layers of secondary deposit.

In the cuticle of the grasses and the mare’s-tails is deposited a large amount of pure flint. So plentiful is this substance, and so equally is it distributed, that it can be separated by heat or acids from the vegetable parts of the plant, and will still preserve the form of the original cuticle, with its cell-walls, stomata, and hairs perfectly well defined.

Fig.7, PlateII., represents a piece of wheat chaff, or “bran,” that has been kept at a white heat for some time, and then mounted in Canada balsam. I prepared the specimen from which the drawing was made by laying the chaff on a piece of platinum, and holding it over the spirit-lamp. A good example of the silex or flint in wheat is often given by the remains of a straw fire, where the stems may be seen still retaining their tubular form but fused together into a hard glassy mass. It is this substance that cuts the fingers of those who handle the wild grasses too roughly, the edges of the blades being serrated with flinty teeth, just like the obsidian swords of the ancient Mexicans, or the shark’s-tooth falchion of the New Zealander.

These are but short and meagre accounts of a very few objects, but space will not permit of further elucidation, and the purpose of this little work is not to exhaust the subjects of which it treats, but to incite the reader to undertake investigation on his own account, and to make his task easier than if he had done it unaided.


                                                                                                                                                                                                                                                                                                           

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