We have hitherto examined what is called the ultimate division of plants. That is, we have looked at each one of the elements separately, and considered its use in vegetable growth.

We will now examine another division of plants, called their proximate division. We know that plants consist of various substances, such as wood, gum, starch, oil, etc., and on examination we shall discover that these substances are composed of the various organic and inorganic ingredients described in the preceding chapters. They are made up almost entirely of organic matter, but their ashy parts, though very small, are (as we shall soon see) sometimes of great importance.

These compounds are called proximate principles,[G] or vegetable proximates. They may be divided into two classes.

The first class are composed of carbon, hydrogen, and oxygen.

The second class contain the same substances and nitrogen.

The first class (those compounds not containing nitrogen) comprise the wood, starch, gum, sugar, and fatty matter which constitute the greater part of all plants, also the acids which are found in sour fruits, etc. Various as are all of these things in their characters, they are entirely composed of the same ingredients (carbon, hydrogen and oxygen), and usually combined in about the same proportion. There may be a slight difference in the composition of their ashes, but the organic part is much the same in every case, so much so, that they can often be artificially changed from one to the other.

As an instance of this, it may be recollected by those who attended the Fair of the American Institute, in 1834, that Prof. Mapes exhibited samples of excellent sugar made from the juice of the cornstalk, starch, linen, and woody fibre.

The ease with which these proximates may be changed from one to the other is their most important agricultural feature, and should be clearly understood before proceeding farther. It is one of the fundamental principles on which the growth of both vegetables depends.

The proximates of the first class constitute usually the greater part of all plants, and they are readily formed from the carbonic acid and water which in nature are so plentifully supplied.

The second class of proximates, though forming only a small part of the plant, are of the greatest importance to the farmer, being the ones from which animal muscle[H] is made. They consist, as will be recollected, of carbon, hydrogen, oxygen and nitrogen, or of all of the organic elements of plants. They are all of much the same character, though each kind of plant has its peculiar form of this substance, which is known under the general name of protein.

The protein of wheat is called gluten—that of Indian corn is zein—that of beans and peas is legumin. In other plants the protein substances are vegetable albumen, casein, etc.

Gluten absorbs large quantities of water, which causes it to swell to a great size, and become full of holes. Flour which contains much gluten, makes light, porous bread, and is preferred by bakers, because it absorbs so large an amount of water.

The protein substances are necessary to animal and vegetable life, and none of our cultivated plants will attain maturity (complete their growth), unless allowed the materials required for forming this constituent. To furnish this condition is the object of nitrogen given to plants as manure. If no nitrogen is supplied the protein substances cannot be formed, and the plant must cease to grow.

When on the contrary ammonia is given to the soil (by rains or otherwise), it furnishes nitrogen, while the carbonic acid and water yield the other constituents of protein, and a healthy growth continues, provided that the soil contains the mineral matters required in the formation of the ash, in a condition to be useful.

The wisdom of this provision is evident when we recollect that the protein substances are necessary to the formation of muscle in animals, for if plants were allowed to complete their growth without a supply of this ingredient, our grain and hay might not be sufficiently well supplied with it to keep our oxen and horses in working condition, while under the existing law plants must be of nearly a uniform quality (in this respect), and if a field is short of nitrogen, its crop will not be large, and of a very poor quality, but the soil will produce good plants as long as the nitrogen lasts, and then the growth must cease.[I]

ANIMALS.

That this principle may be clearly understood, it may be well to explain more fully the application of the proximate constituents of plants in feeding animals.

Animals are composed (like plants) of organic and inorganic matter, and every thing necessary to build them up exists in plants. It seems to be the office of the vegetable world to prepare the gases in the atmosphere, and the minerals in the earth for the uses of animal life, and to effect this plants put these gases and minerals together in the form of the various proximates (or compound substances) which we have just described.

In animals the compounds containing no nitrogen comprise the fatty substances, parts of the blood, etc., while the protein compound, or those which do contain nitrogen, form the muscle, a part of the bones, the hair, and other portions of the animal.

Animals contain a larger proportion of inorganic matter than plants do. Bones contain a large quantity of phosphate of lime, and we find other inorganic materials performing important offices in the system.

In order that animals may be perfectly developed, they must of course receive as food all of the materials required to form their bodies. They cannot live if fed entirely on one ingredient. Thus, if starch alone be eaten by the animal, he might become fat, but his strength would soon fail, because his food contains nothing to keep up the vigor of his muscles. If on the contrary the food of an animal consisted entirely of gluten, he might be very strong from a superior development of muscle, but would not be fat. Hence we see that in order to keep up the proper proportion of both fat and muscle in our animals (or in ourselves), the food must be such as contains a proper proportion of the two kinds of proximates.

It is for this reason that grain, such as wheat for instance, is so good for food. It contains both classes of proximates, and furnishes material for the formation of both fat and muscle. The value of flour depends very much on the manner in which it is manufactured. This will be soon explained.

Apart from the relations between the proximate principles of plants, and those of animals, there exists an important relation between their ashy or inorganic parts; and, food in order to satisfy the demands of animal life, must contain the mineral matter required for the purposes of that life. Take bones for instance. If phosphate of lime is not always supplied in sufficient quantities by food, animals are prevented from the formation of healthy bones. This is particularly to be noticed in teeth. Where food is deficient of phosphate of lime, we see poor teeth as a result. Some physicians have supposed that one of the causes of consumption is the deficiency of phosphate of lime in food.

The first class of proximates (starch, sugar, gum, etc.), perform an important office in the animal economy aside from their use in making fat. They constitute the fuel which supplies the animal's fire, and gives him his heat. The lungs of men and other animals may be called delicate stoves, which supply the whole body with heat. But let us explain this matter more fully. If wood, starch, gum, or sugar, be burned in a stove, they produce heat. These substances consist, as will be recollected, of carbon, hydrogen, and oxygen, and when they are destroyed in any way (provided they be exposed to the atmosphere), the hydrogen and oxygen unite and form water, and the carbon unites with the oxygen of the air and forms carbonic acid, as was explained in a preceding chapter. This process is always accompanied by the liberation of heat, and the intensity of this heat depends on the time occupied in its production. In the case of decay, the chemical changes take place so slowly that the heat, being conducted away as soon as formed, is not perceptible to our senses. In combustion (or burning) the same changes take place with much greater rapidity, and the same amount of heat being concentrated, or brought out in a far shorter time, it becomes intense, and therefore apparent. In the lungs of animals the same law holds true. The blood contains matters belonging to this carbonaceous class, and they undergo in the lungs the changes which have been described under the head of combustion and decay. Their hydrogen and oxygen unite, and form the moisture of the breath, while their carbon is combined with the oxygen of the air drawn into the lungs, and is thrown out as carbonic acid. The same consequence—heat—results in this, as in the other cases, and this heat is produced with sufficient rapidity for the animal necessities. When an animal exercises violently, his blood circulates with increased rapidity, thus carrying carbon more rapidly to the lungs. The breath also becomes quicker, thus supplying increased quantities of oxygen. In this way the decomposition becomes more rapid, and the animal is heated in proportion.

Thus we see that food has another function besides that of forming animal matter, namely to supply heat. When the food does not contain a sufficient quantity of starch, sugar, etc., to answer the demands of the system the animal's own fat is carried to the lungs, and there used in the production of heat. This important fact will be referred to again.