CHAPTER V.

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Ultimate elements of which the body is composed—Proximate principles—Fluids and solids—Primary tissues—Combinations—Results—Organs, systems, apparatus—Form of the body—Division into head, trunk, and extremities—Structure and function of each—Regions—Seats of the more important internal organs.

1. The ultimate elements of which the human body is composed are azote, oxygen, and hydrogen (gaseous fluids); and carbon, phosphorus, calcium, sulphur, sodium, potassium, magnesium, and iron (solid substances). These bodies are called elementary and ultimate, because they are capable of being resolved by no known process into more simple substances.

2. These elementary bodies unite with each other in different proportions, and thus form compound substances. A certain proportion of azote uniting with a certain proportion of oxygen, hydrogen, and carbon, forms a compound substance possessing certain properties. Another proportion of azote uniting with a different proportion of oxygen, hydrogen, and carbon, forms another compound substance possessing properties different from the former. Oxygen, hydrogen, and carbon, uniting in still different proportions without any admixture of azote, form a third compound possessing properties different from either of the preceding. The compounds thus formed by the primary combinations of the elementary substances with each other are called PROXIMATE PRINCIPLES.

3. Each proximate principle constitutes a distinct form of animal matter, of which the most important are named gelatin, albumen, fibrin, oily or fatty matter, mucus, urea, pichromel, osmazome, resin, and sugar.

4. By chemical analysis it is ascertained that all the proximate principles of the body, however they may differ from each other in appearance and in properties, are composed of the same ultimate elements. Gelatin, for example, consists (in 100 parts) of azote 16-988/1000, oxygen 27-207/1000, hydrogen 7-914/1000, carbon 47-881/1000 parts. The elementary bodies uniting in the above proportions form an animal substance, soft, tremulous, solid, soluble in water, especially when heated, and on cooling, which may be considered as its distinctive property, separating from its solution in water into the same solid substance, without undergoing any change in its chemical constitution.

5. Again, albumen consists of azote 15-705/1000, oxygen 23-872/1000, hydrogen 7-540/1000, carbon 52-888/1000, parts. The elementary bodies uniting in these different proportions, there results a second proximate principle, an adhesive fluid, transparent, destitute of smell and taste, miscible in water, but when subjected to a temperature of about 165°, converted into a solid substance no longer capable of being dissolved in water. This conversion of albumen from a fluid, which is its natural state, into a solid, by the application of heat, is called coagulation. It is a process familiar to every one. The white of egg is nearly pure albumen, naturally a glary and adhesive fluid: by boiling, it is coagulated into a white and firm solid.

6. In like manner, fibrin consists of azote 19-934/1000, oxygen 19-685/1000, hydrogen 7-021/1000, carbon 53-360/1000 parts, forming a solid substance of a pale whitish colour and firm consistence, the peculiar character of which is its disposition to arrange itself into minute threads or fibres.

7. On the other hand, fat or oil, which is a fluid substance of a whitish yellow colour, inodorous, nearly insipid, unctuous, insoluble in water and burning with rapidity, consists of a larger proportion of hydrogen, a small proportion of oxygen, and a still smaller proportion of carbon, without any admixture of azote.

8. From this account of the composition of the proximate principles, which it is not necessary to extend further, it is manifest that all of them consist of the same ultimate elements, and that they derive their different properties from the different proportions in which their elements are combined.

9. The ultimate elements that compose the body are never found in a separate or gaseous state, but always in combination in the form of one or other of the proximate principles.

10. In like manner, the proximate principles never exist in a distinct and pure state, but each is combined with one or more of the others. No part consists wholly of pure albumen, gelatin, or mucus, but albumen is mixed with gelatin, or both with mucus.

11. Simple or combined, every proximate principle assumes the form either of a fluid or of a solid, and hence the most general and obvious division of the body is into fluids and solids. But the terms fluid and solid are relative, not positive; they merely express the fact that some of the substances in the body are soft and liquid compared with others which are fixed and hard; for there is no fluid, however thin, which does not hold in solution some solid matter, and no solid, however dense, which does not contain some fluid.

12. Fluids and solids are essentially the same in nature; they differ merely in their mode of aggregation; hence the easy and rapid transition from the one to the other which incessantly takes place in the living body, in which no fluid long remains a fluid, and no solid a solid, but the fluid is constantly passing into the solid and the solid into the fluid.

13. The relative proportion of the fluids in the human body is always much greater than that of the solids; hence its soft consistence and rounded form. The excess, according to the lowest estimate, is as 6 to 1, and according to the highest, as 10 to 1. But the proportion is never constant; it varies according to age and to the state of the health. The younger the age, the greater the preponderance of the fluids. The human embryo, when first perceptible, is almost wholly fluid: solid substances are gradually but slowly superadded, and even after birth the preponderance is strictly according to age; for in the infant, the fluids abound more than in the child; in the child, more than in the youth; in the youth, more than in the adolescent; in the adolescent, more than in the adult; and in the adult, more than in the aged. Thus, among the changes that take place in the physical constitution of the body in the progress of life, one of the most remarkable is the successive increase in the proportion of its solid matter: hence the softness and roundness of the body in youth; its hard, unequal, and angular surface in advanced life; its progressively increasing fixedness and immobility in old age, and ultimate inevitable death.

14. The fluids are not only more abundant than the solids, but they are also more important, as they afford the immediate material of the organization of the body; the media by which both its composition and its decomposition are effected. They bear nourishment to every part, and by them are carried out of the system its noxious and useless matter. In the brain they lay down the soft and delicate cerebral substance; in the bone, the hard and compact osseous matter; and the worn-out particles of both are removed by their instrumentality. Every part of the body is a laboratory in which complicated and transforming changes go on every instant; the fluids are the materials on which these changes are wrought; chemistry is the agent by which they are effected, and life is the governing power under whose control they take place.

15. The fluids, composed principally of water holding solid matter in solution, or in a state of mechanical division, either contribute to the formation of the blood, or constitute the blood, or are derived from the blood; and after having served some special office in a particular part of the system, are returned to the blood; and according to the nature and proportion of the substances they contain, are either aqueous, albuminous, mucous, gelatinous, fibrinous, oleaginous, resinous, or saline.

16. When the analysis of the different kinds of animal matter that enter into the composition of the body has been carried to its ultimate point, it appears to be resolvable into two primitive forms: first, a substance capable of coagulation, but possessing no determinate figure; and secondly, a substance having a determinate figure and consisting of rounded particles. The coagulable substance is capable of existing by itself; the rounded particles are never found alone, but are invariably combined with coagulated or coagulable matter. Alone or combined with the rounded particles, the coagulable matter forms, when liquid, the fluids, when coagulated, the solids.

17. When solid, the coagulable substance is disposed in one of two forms, either in that of minute threads or fibres, or in that of minute plates or laminÆ; hence every solid of the body is said to be either fibrous or laminated. The fibres or laminÆ are variously interwoven and interlaced, so as to form a net-work or mesh; and the interspaces between the fibres or laminÆ are commonly denominated areolÆ or cells (fig. XVII).

18. This concrete substance, fibrous or laminated, is variously modified either alone or in combination with the rounded particles. These different modifications and combinations constitute different kinds of organic substance. When so distinct as obviously to possess a peculiar structure and peculiar properties, each of these modifications is considered as a separate form of organized matter, and is called a PRIMARY TISSUE. Anatomists and physiologists have been at great pains to discriminate and classify these primary tissues; for it is found that when employed in the composition of the body, each preserves its peculiar structure and properties wherever placed, however combined, and to whatever purpose applied, undergoing only such modification as its local connexions and specific uses render indispensable. Considering every substance employed in the construction of the body, not very obviously alike, as a distinct form of organized matter, these primary tissues may be said to consist of five, namely, the membranous, the cartilaginous, the osseous, the muscular, and the nervous.

19. The first primary tissue is the peculiar substance termed MEMBRANE. It has been already stated (16) that one of the ultimate forms of animal matter is a coagulable substance, becoming concrete or solid under the process of coagulation. The commencement of organization seems to be the arrangement of this concrete matter into straight thready lines, at first so small as to be imperceptible to the naked eye. Vast numbers of these threads successively uniting, at length form a single thread of sufficient magnitude to be visible, but still smaller than the finest thread of the silkworm. If the length of these threads be greater than their breadth, they are called fibres; if, on the contrary, their breadth exceed their length, they are termed plates or laminÆ. By the approximation of these fibres or plates in every possible direction, and by their accumulation, combination, and condensation, is constituted the simplest form of organized substance, the primary tissue called membrane.

20. Membrane once formed is extensively employed in the composition of the body: it is indeed the material principally used in producing, covering, containing, protecting, and fixing every other component part of it. It forms the main bulk of the cartilaginous tissue; it receives into its cells the earthy matter on which depend the strength and hardness of the osseous tissue; it composes the canals or sheaths in which are deposited the delicate substance of the muscular, and the still more tender pulp of the nervous tissue; it gives an external covering to the entire body; it lines all its internal surfaces; it envelopes all internal organs; it enters largely as a component element into the substance of every organ of every kind; it almost wholly constitutes all the internal pouches and sacs, such as the stomach, the intestines, the bladder; and all tubes and vessels, such as arteries, veins, and lymphatics; it furnishes the common substance in which all the parts of the body are, as it were, packed; it fills up the interstices between them; it fixes them in their several situations; it connects them all together; in a word, it forms the basis upon which the other parts are superinduced; or rather the mould into which their particles are deposited; so that were it possible to remove every other kind of matter, and to leave this primary tissue unaltered in figure and undiminished in bulk, the general form and outline of the body, as well as the form and outline of all its individual parts, would remain unchanged.

21. The properties which belong to membrane are cohesion, flexibility, extensibility, and elasticity. By its property of cohesion, the several parts of the body are held together; by its combined properties of cohesion, flexibility, and extensibility, the body in general is rendered strong, light, and yielding, while particular parts of it are made capable of free motion. But elasticity, that property by which parts removed from their situation in the necessary actions of life are restored to their natural position, may be regarded as its specific property. The varied purposes accomplished in the economy by the property of elasticity will be apparent as we advance in our subject. Meantime, it will suffice to observe that it is indispensable to the action of the artery in the function of the circulation; to the action of the thorax in the function of respiration; to the action of the joints in the function of locomotion: in a word, to the working of the entire mechanism by which motion of every kind and degree is effected. All these properties are physical, not vital; vital properties do belong even to this primary form of animal matter; but they are comparatively obscure. In the tissue with which organization commences, and which is the least removed from an inorganic substance, the properties that are prominent and essential are merely physical.

22. By chemical analysis, membrane is found to contain but a small proportion of azote, the peculiar element of animal matter. Its proximate principles are gelatin, albumen, and mucus. In infancy and youth, gelatin is the most abundant ingredient; at a more advanced period, albumen predominates[3]. Gelatin differs from albumen in containing a less proportion of azote and a greater proportion of oxygen; on both accounts it must be regarded as less animalized. Thus animalization bears a certain relation to organization. The simplest animal tissue is the least animalized, and the least of all at the earliest period of life. Not only are the physical and mental powers less developed in the young than in the adult, but the very chemical composition of the primary tissue of which the body is constructed is less characteristic of the perfect animal.

23. Membrane exists under several distinct forms; a knowledge of the peculiarities of which will materially assist us in understanding the composition of the body. The simplest form of membrane, and that which is conceived to constitute the original structure from which all the others art produced, is termed the cellular. When in thin slices, cellular membrane appears as a semi-transparent and colourless substance; when examined in thicker masses, it is of a whitish or greyish colour. It consists of minute threads, which cross each other in every possible direction, leaving spaces between them, and thus forming a mesh or net-work (fig. XVII.), not unlike the spider's web. The term cells, given to these interspaces, is employed rather in a figurative sense than as the expression of the fact; for there are no such distinct partitions as the term cell implies. The best conception that can be formed of the arrangement of the component parts of this structure is, to suppose a substance consisting of an infinite number of slender thready lines crossing each other in every possible direction (fig. XVII.). The interspaces between these lines during life, and in the state of health, are filled with a thin exhalation of an aqueous nature, a vapour rather than a fluid, rendering and keeping the tissue always moist. This vapour consists of the thinner part of the blood, poured into these interstitial spaces by a process hereafter to be described, termed secretion. When occupying those spaces, it makes no long abode within them, but is speedily removed by the process of absorption. In health, these two operations exactly equal each other; but if any cause arise to disturb the equilibrium, the vapour accumulates, condenses and forms an aqueous fluid, which distends the cells and gravitates to the most depending parts. Slightly organized as this tissue is, and indistinct as its vita functions may be, it is obvious that it must be the seat of at least two vital functions, secretion and absorption.

Fig. XVII.
A single film of the cellular tissue lifted up and
slightly distended.

24. It is certain that the interspaces or cells of this membrane have no determinate form or size, that they communicate freely with each other, and that this communication extends over the whole body; for if a limb which has been infiltrated be frozen, a thousand small icicles will be formed, assuming the shape of the containing cells, some of which are found to be circular and others cylindrical, and so on. If air or water escape into any particular part of the body, it is often effused over the whole extent of it, and butchers are observed to inflate animals by making a puncture in some part where the cellular tissue is loose, and from this one aperture the air is forced to the most distant parts of the body.

25. Cellular membrane, variously modified and disposed, forms the main bulk of all the other solid parts of the body, constituting their common envelope and bond of union, and filling up all their interstices. It is dense or loose, coarse or fine, according to its situation and office. Wherever it is subject to pressure, it is dense and firm, as in the palm of the hand and the sole of the foot; around the internal organs it is more loose and delicate, and it becomes finer and finer as it divides and subdivides, in order to envelope the soft and tender structures of the body.

Fig. XVIII.
A portion of cellular tissue, very highly magnified, showing
the strings of globules of which its ultimate fibres are by
some supposed to consist.

26. According to some who have carefully examined with the microscope its component threads, they consist of minute particles of a globular figure (fig. XVIII.); other microscopical observers regard the cellular threads as coagulated or condensed animal substance, perfectly amorphous (without form).

27. Every part of this tissue is penetrated by arteries, veins, absorbents, and nerves, endowing it with properties truly vital, though in a less degree than any of the other primary tissues; and varied and important as the uses are which it serves in the economy, the most manifest, though certainly not the only ones, are those which depend upon its physical properties of cohesion, flexibility, extensibility, and elasticity.

Fig. XIX.
1, A portion of adipose tissue; 2, minute bags containing
the fat; 3, a cluster of the bags, separated and suspended.

28. The tissue which contains the fat, termed the adipose, is the second form of membrane; it is obviously a modification of the cellular, from which it differs both in the magnitude of its fibres, whence it constitutes a tougher and coarser web, and in their arrangement; for it is so disposed as to form distinct bags in which the fat is contained. Adipose tissue consists of rounded packets, separated from each other by furrows (fig. XIX. 2, 2); each packet is composed of small spheroidal particles (fig. XIX. 2, 2); each particle is again divisible into still smaller grains, which, on minute inspection, present the appearance of vesicles filled with the adipose matter (fig. XIX. 3).

29. The cells of the cellular tissue, as has been shown (24), are continuous over the whole body; but each adipose vesicle is a distinct bag, having no communication whatever with any other (fig. XIX. 2, 2). The cellular tissue is universally diffused; but the adipose is placed only in particular parts of the body; principally beneath the skin, and more especially between the skin and the abdominal muscles, and around some of the organs contained in the chest and abdomen, as the heart, the kidneys, the mesentery, and the omenta. In most of these situations some portion of it is generally found, whatever be the degree of leanness to which the body may be reduced; while in the cranium, the brain, the eye, the ear, the nose, and several other organs, there is none, whatever be the degree of corpulency. The uses of the fat, which are various, will be stated hereafter.

30. The third form of membrane is termed the

serous. Like the adipose, serous membrane is a modification of the cellular, and, like it also, it is limited in its situation to particular parts of the body, that is, to its three great cavities, namely, the head, the chest, and the abdomen. To the two latter it affords an internal lining, and to all the organs contained in all the three cavities, it affords a covering. By its external surface it is united to the wall of the cavity or the substance of the organ it invests; by its internal surface it is free and unattached; whence this surface is in contact only with itself, forming a close cavity or shut sac, having no communication with the external air. Smooth and polished (fig. XX.), it is rendered moist by a fluid which is supposed to be exhaled in a gaseous state from the serum of the blood; and from this serous fluid the membrane derives its name.

Fig. XX.
A portion of intestine, showing its external surface or
serous coat.

31. Though thin, serous membrane is dense, compact, and of great strength in proportion to its bulk: it is extensible and elastic; extensible, for it expands with the dilatation of the chest in inspiration; elastic, for it contracts with the diminished size of the chest in expiration. In like manner, it stretches with the enlargement of the stomach during a hearty meal, and contracts as the stomach gradually diminishes on emptying itself of its contents. It is furnished with no blood-vessels large enough to admit the colouring matter of the blood; but it is supplied with a great number of the colourless vessels termed exhalents, with the vessels termed absorbents, and with a few nerves. It indicates no vital properties, but those which are common to the simple form of the primary tissue. Its specific uses are to afford a lining to the internal cavities; to furnish a covering to the internal organs; by its polished and smooth surface, to allow a free motion of those organs on each other, and by the moisture with which it is lubricated, to prevent them from adhering together, however closely, or for however long a period they may be in contact.

32. The fourth form of membrane, the fibrous, named from the obvious arrangement of its component parts, consists of longitudinal fibres, large enough to be visible to the naked eye, placed parallel to each other, and closely united. Sometimes these fibres are combined in such a manner as to form a continuous and extended surface, constituting a thin, smooth, dense, and strong membrane, such as that which lines the external surface of bones termed PERIOSTEUM, or the internal surface of the skull (dura mater). At other times, they form a firm and tough expansion (aponeurosis) which descends between certain muscles, separating them from each other, and affording a fixed point for the origin or insertion of neighbouring muscles; or which is stretched over muscles, and sometimes over even an entire limb, in order to confine the muscles firmly in their situation, and to aid and direct their action (fig. XXVII.). Fibrous membrane also constitutes the compact, strong, tough, and flexible bands used for tying parts firmly together, termed LIGAMENTS, principally employed in connecting the bones with each other, and particularly about the joints; and lastly, fibrous membrane forms the rounded white cords in which muscles often terminate, called TENDONS (fig. XXV., XXVI.), the principal use of which is to connect the muscles with the bones, and to serve as cords or ropes to transmit the action of the muscle to a distant point, in the accomplishment of which purposes their operation appears to be entirely mechanical.

33. The fifth form of membrane, the mucous (fig. XXI.), derives its name from the peculiar fluid with which its surface is covered, called mucus, and which is secreted by numerous minute glands, imbedded in the substance of the membrane. As serous membrane forms a shut sac, completely excluding the air, mucous membrane, on the contrary, lines the various cavities which are exposed to the air, such as the mouth, the nostrils, the wind-pipe, the gullet, the stomach, the intestines, the urinary organs, and the uterine system. Its internal surface, or that by which it is attached to the passages it lines, is smooth and dense; its external surface, or that which is exposed to the contact of the air, is soft and pulpy, like the pile of velvet (fig. XXI.). It bears a considerable resemblance to the external surface of the rind of the ripe peach.

Fig. XXI
A portion of the stomach, showing its internal surface
or mucous coat.

Unlike all the other tissues of this class, the mucous membranes are the immediate seat of some of the most important functions of the economy; in the lung, of respiration; in the stomach, of digestion; in one part of the intestine, of chylification; in another, of excretion; while in the mouth and nose, they are the seat of the animal functions of taste and smell; and they are highly organized in accordance with the importance of the functions they perform.

34. The last form of membrane which it is necessary to our present purpose to particularize, is that which constitutes the external covering of the body, and which is called the skin. The skin is everywhere directly continuous with the mucous membranes that line the internal passages, and its structure is perfectly analogous. Both the external and the internal surface of the body may be said therefore to be covered by a continuous membrane, possessing essentially the same organization, and almost identically the same chemical composition. The skin is an organ which performs exceedingly varied and important functions in the economy, to the understanding of which it is necessary to have a clear conception of its structure; some further account of it will therefore be required; but this will be more advantageously given when the offices it serves are explained.

Fig. XXII.
Portions of cartilage, seen in section.

35. Such is the structure, and such are the properties, of the first distinct form of organized matter. The second primary tissue, termed the CARTILAGINOUS (fig. XXII.), is a substance intermediate between membrane and bone. The nature of its organization is not clearly ascertained. By some anatomists, it is regarded as a uniform and homogeneous substance, like firm jelly, without fibres, plates, or cells; others state that they have been able to detect in it longitudinal fibres, interlaced by other fibres in an oblique and transverse direction, but without determinate order. All are agreed that it is without visible vessels or nerves: not that it is supposed to be destitute of them, but that they are so minute as to elude observation. Its manifest properties are wholly mechanical. It is dense, strong, inextensible, flexible, and highly elastic. It is chiefly by its property of elasticity that it accomplishes the various purposes it serves in the economy. It is placed at the extremities of bones, especially about the joints, where, by its smooth surface, it facilitates motion, and, by its yielding nature, prevents the shock or jar which would be produced were the same kind and degree of motion effected by a rigid and inflexible substance. Where a certain degree of strength with a considerable degree of flexibility are required, it supplies the place of bone, as in the spinal column, the ribs and the larynx.

Fig. XXIII.
Membranous portion of bone; the osseous portion being
so completely removed, that the bone is capable of being tied in a knot.

36. The third distinct form of organized matter is termed the OSSEOUS tissue. Bone is composed of two distinct substances, an animal and an earthy matter: the former organic, the latter inorganic. The animal or organic matter is analogous both in its nature and in its arrangement to cellular tissue; the earthy or inorganic matter consists of phosphoric acid combined with lime, forming phosphate of lime. The cellular tissue is aggregated into plates or laminÆ, which are placed one upon another, leaving between them interspaces or cells, in which is deposited the earthy matter (phosphate of lime). If a bone, for example, the bone called the radius, one of the bones of the fore-arm, be immersed in diluted sulphuric, nitric, muriatic, or acetic acid, it retains its original bulk and shape; it loses, however, a considerable portion of its weight, while it becomes so soft and pliable, that it may be tied in a knot (fig. XXIII.). In this case, its earthy matter is removed by the agency of the acid, and is held in solution in the fluid; what remains is membranous matter (cellular tissue). If the same bone be placed in a charcoal fire, and the heat be gradually raised to whiteness, it appears on cooling as white as chalk; it is extremely brittle; it has lost much of its weight, yet its bulk and shape continue but little changed. In this case, the membraneous matter is wholly consumed by the fire, while the earth is left unchanged (fig. XXIV.). Every constituent atom of bone consists, then, essentially of animal and earthy matter intimately combined. A little more than one-third part consists of animal matter (albumen), the remaining two-thirds consist of earthy matter (phosphate of lime); other saline substances, as the fluate of lime and the phosphate of magnesia, are also found in minute quantity, but they are not peculiar to bone.

Fig. XXIV.
Earthy portion of bone.

37. In general, the osseous tissue is placed in the interior of the body. Even when bone approaches the surface, it is always covered by soft parts. It is supplied with but few blood-vessels, with still fewer nerves, with no absorbents large enough to be visible, so that though it be truly alive, yet its vital properties are not greatly developed. The arrangement of its component particles is highly curious; the structure, the disposition, and the connexion of individual bones afford striking examples of mechanism, and accomplish most important uses in the economy; but those uses are dependent rather upon mechanical than vital properties. The chief uses of bone are— 1. By its hardness and firmness to afford a support to the soft parts, forming pillars to which the more delicate and flexible organs are attached and kept in their relative positions. 2. To defend the soft and tender organs by forming a case in which they are lodged and protected, as that formed by the bones of the cranium for the lodgment and protection of the brain (fig. XLVII.); by the bones of the spinal column for the lodgment and protection of the spinal cord (fig. XLVIII.); by the bones of the thorax (fig. LIX.), for the lodgment and protection of the lungs, the heart, and the great vessels connected with it (fig. LIX.). 3. By affording fixed points for the action of the muscles, and by assisting in the formation of joints to aid the muscles in accomplishing the function of locomotion.

38. All the primary tissues which have now been considered consist of precisely the same proximate principles. Albumen is the basis of them all; with the albumen is always mixed more or less gelatin, together with a minute quantity of saline substance: to the osseous tissue is superadded a large proportion of earthy matter. With the exception of the mucous, the organization of all these tissues is simple; their vital properties are low in kind and in degree; their decided properties are physical, and the uses they serve in the economy are almost wholly mechanical.

Fig. XXV.
Portion of a muscle; showing (a) the muscular fibres
and their parallel direction; and (b) the termination of
the fibres in tendon.

39. But we next come to a tissue widely different in every one of those circumstances, a tissue consisting of a new kind of animal matter, and endowed with a property not only peculiar to itself, but proper to living substance, and characteristic of a high degree of vital power. Muscular tissue, the fourth distinct form of animal matter, commonly known under the name of flesh, is a substance resembling no other in nature. It consists of a soft and pulpy substance, having little cohesive power, arranged into fibres which are distinctly visible to the naked eye, and which are disposed in a regular and uniform manner, being placed close and parallel to each other (fig. XXV.). These fibres are every where pretty uniformly the same in shape, size, and general appearance, being delicate, soft, flattened, and though consisting only of a tender pulp, still solid (fig. XXV.). When examined under the microscope, fibres, which to the naked eye appear to be single threads, are seen to divide successively into smaller threads, the minutest or the ultimate division not exceeding, as is supposed, the 40,000th part of an inch in diameter. On the other hand, the fibres which are large enough to be visible to the naked eye, are obviously aggregated into bundles of different magnitude in different muscles, but always of the same uniform size in the same muscle (fig. XXV.).

Fig. XXVI.
Two portions of muscle; one of which, a, is covered with
membrane; the other, b, is uncovered; c, the muscular
fibres terminating in tendon.

40. The ultimate thread, or the minutest division of which the muscular fibre is susceptible, is called a filament; the smallest thread which can be distinguished by the naked eye is termed a fibre (fig. XXVI.); and the bundle which is formed by the union of fibres is denominated a fasciculus. The proper muscular substance is thus arranged into three distinct forms progressively increasing in size,—the filament, the fibre, and the fasciculus. The filament, the fibre, the fasciculus, as well as the muscle itself, formed by the aggregation of fasciculi, is each inclosed in its own distinct sheath of cellular membrane (fig. XXVI. a).

Fig. XXVII.
Portion of a muscle enclosed in a sheath of fascia
or aponeurosis.

41. The composition of the ultimate filament has been very carefully examined by many distinguished physiologists with microscopes of high magnifying power. Under some of these microscopes the filament appears to consist of a series of rounded particles or globules of the same size as the particles of the blood when deprived of their colouring matter, so that it looks like a string of pearls (fig. XXVIII.), each globule being commonly stated to be about the 2000th part of an inch in diameter. But it is now pretty generally agreed that this globular appearance of the ultimate muscular fibre vanishes under the more improved microscopes of the present day, and, as viewed by the latter, appears as a peculiar pulpy substance arranged into threads of extreme minuteness, placed close and parallel to each other, intersected by a great number of delicate lines passing transversely across the muscular threads (fig. XXIX.),

Fig. XXVIII.
Ultimate fibres of muscle, very greatly magnified; showing
the strings of globules of which they are supposed by
some to consist.

42. With the exception of the organs of sense, the muscular tissue is more abundantly supplied with arteries, veins, and nerves, than any other substance of the body. Every ultimate thread or filament appears to be provided with the ultimate branch of an artery, vein, and nerve. These vessels are seen ramifying on the surface of the delicate web of membrane that incloses the pulp, but cannot be traced into it.

Fig. XXIX.
The appearance of the ultimate muscular fibres and of
their transverse lines, as seen under the microscope of Mr. Lister,
when the object is magnified 500 diameters.

43. The proximate principle of which the muscular pulp is composed is fibrin. From the pulp, when inclosed in its sheath of membrane, albumen, jelly, various salts, and a peculiar animal extract called osmazome, are also obtained; but these substances are probably derived from the membranous, not the muscular, matter. Fibrin contains a larger proportion of azote, the element peculiar to the animal body, and by the possession of which its chemical composition is distinguished from that of the vegetable, than any other animal substance.

Fig. XXX.
Portion of the trunk of
a nerve; dividing into branches.

44. Muscular tissue possesses a slight degree of cohesion, a high degree of flexibility and extensibility, but no degree of elasticity; for although muscle, considered as a compound of muscular substance and membrane, be highly elastic, yet this property is probably altogether owing to the membranous matter in which it is enveloped. Its peculiar and distinctive property is vital, not physical, and consists in the power of diminishing its length, or of contracting or shortening itself on the application of a stimulus. This property, which is termed contractility, is the great, if not the sole source of motion in the body. Without doubt, elasticity and gravity, under the generating and controlling powder of contractility, aid in accomplishing various kinds of motion. Thus membranes, tendons, ligaments, cartilages, and bones, by their physical and mechanical properties, modify, economize, facilitate, concentrate and direct the motive power generated by the pure muscular substance; but still the only real source of motion in the body is muscular tissue, and the only mode in which motion is generated is by contractility. This will be more fully understood hereafter.

Fig. XXXI.
Ultimate fibres of nerve, very highly magnified; showing
the strings of globules of which they consist.

45. The last primary tissue, termed the NERVOUS, is equally distinct in nature and peculiar in property. It consists of a soft and pulpy matter, of a brownish white colour (fig. XXX.). According to some, the nervous, like the muscular pulp, is composed of minute globules, arranged in the same manner like a string of pearls (fig. XXXI.); according to others, it consists of solid elongated threads, of a cylindrical form, differing in thickness from that of a hair to the finest fibre of silk. The pulp, whatever its form of aggregation, is inclosed in a sheath of delicate cellular tissue. This external or containing membrane is called the neurilema, or the nerve-coat; the internal or contained substance, the proper nervous matter, is termed the nerve-string. The nerve-string, enveloped in its nerve-coat, constitutes the nervous filament. As in the muscle, so in the nerve, many filaments unite to form a fibre, many fibres to form a fasciculus, and many fasciculi to form the large cord termed a nerve. Moreover, as in the muscle, so in the nerve, the filament, the fibre, the fasciculus, the nervous cord itself, are each enveloped in its own distinct sheath of cellular membrane; but the arrangement of the nervous fibres differs from that of the muscular in this, that though the nervous fibres are placed in juxtaposition, yet they do not, like the muscular, maintain through their entire course a parallel disposition, but cross and penetrate each other, so as to form an intimate interlacement (fig. XXXII.).

Fig. XXXII.
Nervous fibres, deprived of their neurilema and unravelled,
showing the smaller threads, or filaments, of which
the fibres consist.

46. The nervous pulp is at least as liberally supplied with blood-vessels as the muscular; the vessels are spread out upon the nerve-coat, in which they divide into innumerable branches of extreme minuteness, the distribution of which is so perfect, that there is not a particle of nervous matter which is not supplied both with an arterial and a venous vessel. Hence the neurilema is not merely a sheath containing and protecting the nervous pulp, but it affords an extended mechanical surface for sustaining the arterial vessels, from which the pulp is probably secreted, and certainly nourished.

47. Albumen, in conjunction with a peculiar fatty matter, constitutes the chief proximate principles of which the nervous tissue is composed. To these are added a small proportion of the animal substance termed osmazome, a minute quantity of phosphorus, some salts, and a very large proportion of water; for out of one hundred parts of nervous substance, water constitutes as much as eighty. Its peculiar vital property is sensibility; and as all motion depends on the contractility of the muscular fibre, so all sensation depends on the sensibility of the nervous substance.

48. Such are the primary tissues, or the several kinds of organized matter of which the body is composed; and from this account it is obvious that they consist of three only—namely, the concrete matter forming the basis of membrane, the pulpy matter forming the proper muscular substance, and the pulpy matter forming the proper nervous substance. Of these three kinds of animal matter the component parts of the body consist. In combining to form the different structures, these primary substances are intermixed and arranged in a great variety of modes; and from these combinations and arrangements result either an organ, a system, or an apparatus.

49. As filaments unite to form fibres, and fibres to form tissues, so tissues unite to form organs: that is, bodies having a determinate size and figure, and capable of performing specific actions. The cellular, the muscular, and the nervous tissues are not organs; membranes, muscles, and nerves are organs. The tissue, the simple animal substance, is merely one of the elements of which the organ is composed; the organ is compounded of several of those simple substances, arranged in a determinate manner, and moulded into a given shape, and so constituting a specific instrument. The basis of the muscle is muscular tissue; but to this are added, invariably, membrane, often tendon, and always vessels and nerves. It is this combination that forms the specific instrument called a muscle, and that renders it capable of performing its specific action. And every such combination, with its appropriate endowment, constitutes an organ.

50. Organs are arranged into groups or classes, according as they possess an analogous structure, and perform an analogous function; and this assemblage constitutes a SYSTEM. All the muscles of the body, for example, whatever their size, form, situation, or use, have an analogous structure, and perform an analogous function, and hence are classed together under the name of the muscular system. All the bones, whatever their figure, magnitude, density, position, or office, are analogous in structure and function; and hence are classed together under the name of the osseous system. For the same reason, all the cartilages, ligaments, vessels and nerves, form respectively the cartilaginous, ligamentous, vascular and nervous systems.

51. An APPARATUS, on the contrary, is an assemblage of organs, it may be differing widely from each other in structure, and exercising various and even opposite functions; but all nevertheless concurring in the production of some common object. The apparatus of nutrition consists of the organs of mastication, deglutition, digestion, absorption, and assimilation. Among the individual organs which concur in carrying on these functions may be reckoned the lips, the teeth, the tongue, the muscles connected with the jaws, the gullet, the stomach, the duodenum, the small intestines, the pancreas, the liver, the lacteal vessels, the mesenteric glands, and the lungs. Many of these organs have no similarity in structure, and few have any thing analogous in function; yet all concur, each in its appropriate mode and measure, to the conversion of the aliment into blood. In the apparatus of respiration, in that of circulation, of secretion, of excretion; in the apparatus of locomotion, in the apparatus of sensation, and more especially in the apparatus of the specific sensations,—vision, hearing, smell, taste, touch, organs are combined which have nothing in common but their concurrence in the production of a common end: but this concurrence is the principle of their combination; and the individual organs having this conjoint operation, taken together, constitute an apparatus.

52. A clear idea may now be affixed to the terms structure and organization. Structure may be considered as synonymous with arrangement; the disposition of parts in a determinate order; that which is constructed or built up in a definite mode, according to a determinate plan. The arrangement of the threads of the cellular web into areolÆ or cells; the combination of the primary threads into fibres or laminÆ; the disposition of the muscular pulp into filaments, placed parallel to each other; the investment of the filaments in membraneous sheaths; the combination of the filaments, included in their sheaths, into fibres; the aggregation of fibres into fasciculi; and the analogous arrangement and combination of the nervous pulp, are examples of structure. But when those structures are applied to particular uses; when they are so combined and disposed as to form a peculiar instrument, endowed with a specific function; when the cellular fibres, for example, are so arranged as to make a thin, dense, and expanded tissue; when to this tissue are added blood-vessels, absorbents, and nerves; when, in a word, a membrane is constructed, an organ is formed; when, in like manner, to the muscular and the nervous fibres, arranged and moulded in the requisite mode, are added blood-vessels, absorbents, and nerves, other organs are constructed capable of performing specific functions: and this is organization—the building up of organs—the combination of definite structures into special instruments. Structure is the preparatory process of organization; the one is the mere arrangement of the material; the other is the appropriation of the prepared material to a specific use.

53. The term organization is employed in reference both to the component parts of the body, and to the body considered as a whole. We speak of an organized substance and of an organized body. An organized substance is one in which there is not only a definite arrangement of its component parts (structure), but in which the particular arrangement is such as to fit it for accomplishing some special use. Every organized substance is therefore essentially a special organ; limited in its object it may be, and perhaps only conducive to some further object; but still its distinctive character is, that it has a peculiar structure, fitting it for the accomplishment of some appropriate purpose. On the other hand, an organized body is a congeries of organs—the aggregate of the individual organs. Attention was directed in the early part of this work to one peculiar and essential character, by which such an organized is distinguished from an unorganized body. Between the individual parts of the organized body there is so close a relation, that no one of them can be removed or injured, or in any manner affected without a corresponding affection of the whole. The action of the heart cannot cease without the cessation of the action of the lung; nor that of the lung without that of the brain; nor that of the brain without that of the stomach; in a word, there is no organ in whatever distant nook of the system it be placed, or however apparently insignificant its function, that is not necessary to the perfection of the whole. But into whatever number of portions an unorganized body may be divided, each portion retains the properties of the mass, and constitutes in itself a perfect existence; there being no relation between its individual parts, excepting that of physical attraction: on the contrary, each component part of an organized body, being endowed with some appropriate and specific power, on the exercise of which the powers of all the other parts are more or less dependent, the whole must necessarily suffer if but one part fail.

54. From the whole, then, we see that the human body is a congeries of organs; that those organs are constructed of a few simple tissues; and that all its parts, numerous, diversified, and complex as they are, are composed of but three primary forms of animal matter variously modified and combined.

55. But though by the analysis of its component parts, this machine, so complex in its construction, and so wonderfully endowed, may be reduced to this state of simplicity; and although this analytical view of it be highly useful in enabling us to form a clear conception of the nature of its composition; yet it is only by considering its individual parts such as they actually are, and by studying their situation, connexion, structure, and action, that we can understand it as a whole, and apply our knowledge of it to any practical use.

56. Viewing then the human body as a complicated whole, as a congeries of organs made up of various combinations of simple tissues, it may be observed, in reference to its external configuration, that it is rounded. This rounded form is principally owing to the large proportion of fluids which enter into its composition. The roundness of the face, limbs, and entire surface of the child, are in striking contrast to the unequal and irregular surface of the old man, whose humours are comparatively very much smaller in quantity.

57. The length of the human body exceeds its breadth and thickness; the degree of the excess varying at different periods of life, and according to the peculiar constitution of the individual. In the extremities, the bones, muscles, vessels, and nerves, are especially distinguished by their length.

Fig. XXXIV.
Front view of the skeleton. 1. the head; 2. the trunk;
3. the superior extremities; 4. the inferior extremities.

Fig. XXXV.
Back view of the skeleton. 1. the head; 2. the trunk; 3. the superior extremities; 4. the inferior extremities.

58. The form of the human body is symmetrical, that is, it is capable of being divided into two lateral and corresponding halves. Suppose a median line to pass from the vertex of the head through the centre of the spinal column (fig. XXXIV. 1, 2); if the body be well formed, it will be divided by this line into two exactly equal and corresponding portions (fig. XXXV. 1). This symmetrical disposition of the body is not confined to its external configuration. It is true of many of the internal organs; but principally, as has been already stated, of those that belong to the animal life. The brain and the spinal cord are divisible into two exactly equal halves (figs. XLVIII. d, and XLIX. 1, 2, 3); the organs of sense are double and symmetrical: the muscles of one side of the body exactly correspond to those of the other (fig. XXXIII.); the two hands and arms and the two lower extremities are alike (figs. XXXIV., XXXV.); but for the most part, the organs of the organic life, the stomach, the intestines, the liver, the spleen, for example, are single, and not symmetrical.

59. The human body is divided into three great portions, the head, the trunk, and the extremities (figs. XXXIV. and XXXV. 1, 2, 3, 4).

60. By the head is meant all that part of the body which is placed above the first bone of the neck (fig. XXXIV. 1). It is of a spheroidal figure, broader and deeper behind than before, somewhat like an egg in shape, with the broad end behind; it is flattened at its sides (figs. XXXV. 1, and XXXVI. 2, 4). Its peculiar figure renders it at once stronger and more capacious than it could have been had it possessed any other form. It is supported by its base on the spinal column, to which it is attached by the peculiar structure termed a joint (fig. XXXIV.), and fastened by ligaments of exceeding strength.

61. The head contains the central organ of the nervous system; the organs of the senses, with the exception of that of touch; and the organs of mastication. It comprehends the cranium and the face. Both are composed partly of soft parts, as the teguments, namely, skin, fat, &c., and muscles; and partly of bones.

Fig. XXXVI.
1. Frontal bone; 2. parietal bone; 3. occipital bone;
4. temporal bone; 5. nasal bone; 6. malar bone; 7. superior
maxillary bone; 8. inferior maxillary bone.

Fig. XXXVII.
Bones of the skull, separated; front view. 1. Frontal
bone; 2. portions of the parietal bones; 3. malar or cheek
bones; 4. nasal bones; 5. superior maxillary or bones of
the upper jaw; 6. the vomer; 7. the inferior maxillary or
bone of the lower jaw.

Fig. XXXVIII.
Bones of the skull separated; side view. 1. Frontal
bone; 2. parietal bone; 3. occipital bone; 4. temporal
bone; 5. nasal bone; 6. malar bone; 7. superior maxillary
bone; 8. the unguis; 9. the inferior maxillary bone.

Fig. XXXIX.
Bones forming the base of the skull; viewed from the
inside. 1. Occipital bone; 2. temporal bones; 3. sphenoid
bone; 4. ethmoid bone; 5. superior maxillary bones, or
bones of the upper jaw; 6. malar or cheek bones; 7. foramen
magnum.

62. The bones of the cranium are eight in number, six of which are proper to the cranium, and two are common to it and to the face. The six bones proper to the cranium are the frontal (fig. XXXVII. 1), the two parietal (fig. XXXVI. 2), the two temporal (fig. XXXVIII. 4), and the occipital (fig. XXXVIII. 3); the two common to the cranium and face are the ethmoidal (fig. XXXIX. 4), and the sphenoidal (fig. XXXIX.3). The frontal bone forms the entire forepart of the vault (fig. XXXVII. 1); the two parietal form the upper and middle part of it (fig. XXXVIII. 2); the two temporal form the lower part of the sides (fig. XXXVIII. 4); the occipital forms the whole hinder part, together with a portion of the base (figs. XXXVIII. 3, XXXVI. 3, XXXIX. 1); while the ethmoidal forms the forepart, and the sphenoidal the middle part of the base (fig. XXXIX. 3, 4).

Fig. XL.
Portions of the bones of the cranium; showing the corresponding
inequalities in their margins: which margins,
when in apposition, constitute the mode of union termed
suture. 1. External surface of the bone; 2. internal surface.
Fig. XLI and Fig. XLII.
1. Side view of the adult skull, showing the several bones
united by suture; 2. side view of the foetal skull, showing
the bones imperfectly ossified, separated to some extent
from each other, the interspace being occupied by membrane.
The small size of the face compared with that of
the cranium is strikingly apparent.

63. These bones are firmly united together. The union of bones is technically called an articulation or joint. All joints are either immoveable or moveable. The union of the bones of the cranium affords an example of an immoveable articulation. Prominences and indentations, like the teeth of a saw, are formed in the margins of the contiguous bones (figs. XXXVIII. and XL.). At these inequalities of surface, which are exactly adapted to each other (figs. XXXVIII. and XL.), the two bones are in immediate apposition in such a manner as to preclude the possibility of motion, and even to render the separation extremely difficult. This mode of articulation is termed a suture. There are certain advantages in constructing the cranium of several distinct bones, and in uniting them in this peculiar mode. 1. The walls of the vault are stronger than they could have been had they been formed of a single piece. 2. In the foetus, the bones are at some distance from each other (fig. XLII.); at birth, they yield and overlap one another; and in this manner they conduce to the security and ease of that event. 3. Minute vessels pass abundantly and securely through the interstices of the sutures to and from the interior of the cranium; in this manner, a free communication is established between the vessels within and without this cavity. 4. It is probable that the shock produced by external violence is diminished in consequence of the interruption of the vibration occasioned by the suture; it is certain that fracture is prevented by it from extending as far as it would do in one continued bony substance.

Fig. XLIII.
Section of the skull. 1. Cavity of the cranium occupied by
the brain; 2. cut edge of the bones of the cranium, showing
the two tables of compact bone and the intervening spongy
texture called diploË.

64. The vault of the cranium forms a cavity which contains the brain (fig. XLIII.and XLVIII.) The size of this cavity is invariably proportioned to that of the organ it lodges and protects. The form and magnitude of the cavity, and consequently the shape and size of the cranium, depend upon the brain, and not of the brain upon the cranium. The soft parts model and adapt to themselves the hard, and not the hard the soft. The formation of the brain in the foetus is anterior to that of the case which ultimately contains it; and the hard bone is moulded upon the soft pulp, not the pulp upon the bone. At every period of life, on the inner surface of the cranium there are visible impressions made by the convolutions of the brain, and the ramifications of the arteries (figs. XXXIX. 1, 2, and XL. 2), and on its external surface are depressions occasioned by the action of the external muscles. Nor does the modifying power of the brain over the bones of the cranium terminate at birth. The formation of bone, always a slow process, is never completed until the child has attained its third or fourth year, and often not until a much later period. At this tender age, the bones, which in advanced life are hard and rigid, are comparatively soft and yielding, and consequently more readily receive and retain the impression of the convolutions and of the other projecting parts of the brain, by which they are sometimes so deeply marked, that an attentive examination of the inner surface of the cranium is of itself sufficient to determine not only that some part, but to indicate the very part of the brain which has been preternaturally active. At this tender age, pressure, internal or external, general or partial, may readily change the form of the cranium. If, by a particular posture, the head of a child be unequally balanced on the spine, the brain will press more on that side of the cranium than on the other; the organ will expand in the direction to which it inclines; that portion of it will become preternaturally developed, and consequently the balance of its functions will be disturbed. An awkward way of standing or sitting, perhaps contracted inadvertently and kept up by habit; a wry neck; any cause that keeps the head constantly inclined to one side, may produce this result, examples of which and of its consequences will be given hereafter.

65. Tracing them from without inwards we see, then, that the various coverings afforded to the brain, the central organ of the animal life, seated in its vaulted cavity, are: 1. The tegument, consisting of the skin and of cellular and adipose membrane. 2. Beneath the tegument, muscles, in the forepart and at the vertex, comparatively slender and delicate; at the sides and posteriorly, thick, strong, and powerful (fig. XLIV.). 3. Beneath the muscles, a thin but dense membrane, termed the pericranium, lining the external surface of the cranial bones. 4. Beneath the pericranium, the bony substance of the cranium, consisting of two firm and hard bony plates, with a spongy, bony structure, called diploË, interposed between them (fig. XLIII. 2). 5. Immediately in contact with the inner surface of the bony substance of the cranium, and forming its internal lining, the dense and strong membrane, called the dura mater, not only affording a general covering to the brain, but sending firm partitions between individual portions of it (fig. XLVIII. c.). 6. A serous membrane lining the internal surface of the dura mater, and reflected over the entire surface of the brain, termed the arachnoid tunic. 7. A thin and delicate membrane in immediate contact with the substance of the brain, descending between all its convolutions, lining all its cavities and enveloping all its fibres, called the pia mater. 8. An aqueous fluid, contained between the arachnoid membrane and the pia mater. Skin, muscle, pericranium, bone, dura mater, arachnoid membrane, pia mater, and aqueous fluid, superimposed one upon another, form, then, the covering and defence of the brain; so great is the care taken to protect this soft and tender substance.

66. The bones of the face consist of fourteen, namely, the two superior maxillary or jaw-bones (fig. XXXVII. 5), the two malar or cheek bones (fig. XXXVII. 3), the two nasal bones (fig. XXXVII. 4), the two palate bones, the two ossa unguis (fig. XXXVIII. 8), the two inferior turbinated bones, the vomer (fig. XXXVII. 6), and the inferior maxilla or the lower jaw (fig. XXXVII. 7.) This irregular pile of bones is divided into the superior and inferior maxilla or jaws; the superior maxilla being the upper and immoveable portion of the face; the inferior maxilla being the lower and moveable portion of it. Besides these bones, the face contains thirty-two teeth, sixteen in each jaw. The bones of the upper jaw are united together by sutures, and the union is so firm, that they have no motion but what they possess in common with the cranium. The lower jaw is united by a distinct articulation with the cranium (figs. XXXIV. and XXXV.).

67. Besides the bones and the teguments, the face contains a number of muscles, which for the most part are small and delicate (fig. XLIV.), together with a considerable portion of adipose matter; while, as has been stated, the face and head together contain all the senses, with the exception of that of touch, which is diffused, more or less, over the entire surface of the body.

Fig. XLIV.
Muscles of the face.

68. The second great division of the body, termed the TRUNK, extends from the first bone of the neck to that called the pubis in front, and to the lower end of the coccyx behind (fig. XXXIV. 2). It is subdivided into the thorax, the abdomen, and the pelvis (fig. XLV.).

69. The thorax or chest extends above from the first bone of the neck, by which it is connected with the head, to the diaphragm below, by which it is divided from the abdomen (figs. XLV. and LXI.). It consists partly of muscles and partly of bones; the muscular and the osseous portions being in nearly equal proportions. Both together form the walls of a cavity in which are placed the central organs of circulation and respiration (fig. LX. 2, 5). The chief boundaries of the cavity of the thorax before, behind, and at the sides, are osseous (fig. XLV.); being formed before, by the sternum or breast-bone (fig. XLV. 6); behind, by the spinal column or back bone (fig. XLV. 2, 4); and at the sides, by the ribs (fig. XLV. 7). Below, the boundary is muscular, being formed by the diaphragm (fig. LXI. 2), while above the thorax is so much contracted (fig. XLV.), that there is merely a space left for the passage of certain parts which will be noticed immediately.

70. The figure of the thorax is that of a cone, the apex being above (fig. XLV.), through the aperture of which pass the tubes that lead to the lungs and stomach, and the great blood-vessels that go to and from the heart (fig. LX.). The base of the cone is slanting, and is considerably shorter before than behind, like an oblique section of the cone (fig. XLV.).

71. The osseous portion of the walls of the thorax is formed behind by the spinal column, a range of bones common indeed to all the divisions of the trunk; for it constitutes alike the posterior boundary of the thorax, abdomen, and pelvis (fig. XLV. 2, 4, 6). It is composed of thirty distinct bones, twenty-four of which are separate and moveable on one another, and on this account are called true vertebrÆ (fig. XLV. 2, 4); the other five, though separate at an early period of life, are subsequently united into a single solid piece, called the sacrum (fig. XLV. 5). The bones composing this solid piece, as they admit of no motion on each other, are called false vertebrÆ (fig. XLV. 5). To the extremity of the sacrum is attached the last bone of the series, termed the coccyx (fig. XXXV.).

72. From above downwards, that is, from the first bone of the neck to the first bone of the sacrum, the separate bones forming the column progressively increase in size; for this column is the chief support of the weight of the head and trunk, and this weight is progressively augmenting to this point (fig. XLV. 2, 4). From the sacrum to the coccyx, the bones successively diminish in size, until, at the extremity of the coccyx, they come to a point (fig. XXXV.). The spinal column may therefore be said to consist of two pyramids united at their base (fig. XLV. 4, 5). The superior pyramid is equal in length to about one third of the height of the body, and it is this portion of the column only that is moveable.

73. The two surfaces of the spinal column, the anterior and the posterior, present a striking contrast (figs. XXXIV. and XXXV.). The anterior surface, which in its whole extent is rounded and smooth, is broad in the region of the neck, narrow in the region of the back, and again broad in the region of the loins (fig. XLV. 2, 4.). It presents three curvatures (fig. XLV. 2, 4); the convexity of that of the neck being forwards, that of the back backwards, and that of the loins again forwards (fig. XLV. 2, 4).

74. From the posterior surface of the column, which is every where irregular and rough, spring, along the median line, in regular series, strong, sharp, and pointed projections of bone (fig. XXXV.), which from being sharp and pointed, like elongated spines, are called spinous processes, and have given name to the whole chain of bones. These processes afford fixed points for the action of powerful muscles. Extending the whole length of the column, from the base of the skull to the sacrum, on each side of the spinous processes, are deep excavations, which are filled up with the powerful muscles that maintain the trunk of the body erect.

75. From the lateral surfaces of the column likewise spring short but strong projections of bone, termed transverse processes, which also give attachment to powerful muscles (fig. XLVI.).

Fig. XLV.
Bones of the trunk. 1. Spinal column; 2. the seven cervical
vertebrÆ; 3. the twelve dorsal vertebrÆ; 4. the five
lumbar vertebrÆ; 5. the sacrum; 6. the sternum; 7. the
true ribs; 8. the false ribs; 9. the clavicle; 10. the scapula;
11. the ilium; 12. the ischium; 13. the pubes;
14. the acetabulum; 15. the brim of the pelvis.

76. The separate bones of the series have a kind of turning motion on each other; hence each is called a vertebra, and the name of vertebral column is often given to the entire series, as well as that of spinal column. That portion of the column which forms the neck consists of seven distinct bones, called cervical vertebrÆ (fig. XLV. 2); that portion which forms the back consists of twelve, called dorsal vertebrÆ (fig. XLV. 3); that portion which forms the loins consists of five, called lumbar vertebrÆ (fig. XLV. 4). Between each of these classes of vertebrÆ there are specific differences, but they need not be described here: all that is necessary to the present purpose is an account of the structure which is common to every vertebra.

77. By inspecting fig. XLVI. 1, it will be seen that the upper and under edges of each vertebra consist of a ring of bone, of a firm and compact texture, rendering what may be called the body of the vertebra exceedingly strong (fig. XLVI. 3). This ring of bone forms a superficial depression (fig. XLVI. 2), for the reception of a peculiar substance, immediately to be described, which is interposed between each vertebra (fig. XLVII. 2).

78. The anterior surface of the body of the vertebra is convex (fig. XLVI. 3); its posterior surface is concave (fig. XLVI. 4); from the posterior surface springs a bony arch (figs. XLVI. 5 and LIII. 1), which, together with the posterior concavity, forms an aperture of considerable magnitude (fig. XLVI. 6), a portion of the canal for the passage of the spinal cord (figs. XLVII. 3, and XLIX. 3).

Fig. XLVI.
View of some of the vertebrÆ, which by their union form
the spinal column.
a. A vertebra of the neck; b. a vertebra of the back;
a vertebra of the loins.
1. Ring of compact bone forming, 3, the body of the
vertebra; 2. superficial depression for the reception of the
intervertebral cartilage; 3. anterior surface of the body of
the vertebra; 4. posterior surface; 5. bony arch; 6. opening
for the passage of the spinal cord; 7. opening for the passage
of the spinal nerves; 8. articulating processes by
which the vertebrÆ are joined to each other; 9. two dorsal
vertebrÆ united, showing the arrangement of, 10, the spinous
processes; 11. a portion of a rib articulated with the side
of the vertebra.

79. Both the upper and under edges of the arch form a notch (fig. XLVI. 7.), which, together with a corresponding notch in the contiguous vertebra, completes another aperture rounder and smaller than the former, but still of considerable size (fig. XLVI. 7.), the passage of the spinal nerves (fig. XLVII. 3).

80. From both the upper and under sides of the arch proceed two short but strong projections of bone (fig. XLVI. 8.), termed the articulating processes, because it is chiefly by these processes that the vertebrÆ are connected together. From the beginning to the end of the series, the two upper processes of the one vertebra are united with the two lower processes of the vertebra immediately above it (fig. XLVI. 9), and around the edges of all the articulating processes are visible rough lines, which mark the places to which the articulating ligaments are attached.

81. No vertebra, except the first, rests immediately upon its contiguous vertebra (fig. XLV. 2, 4). Each is separated from its fellow by a substance of a peculiar nature interposed between them, termed the intervertebral substance (figs. XLVII. 2, and L. 2). This substance partakes partly of the nature of cartilage, and partly of that of ligament. It is composed of concentric plates, formed of oblique fibres which intersect each other in every direction. This substance, for about a quarter of an inch from its circumference towards its centre, is tough, strong, and unyielding; then it becomes softer, and is manifestly elastic; and so it continues until it approaches the centre, when it becomes pulpy, and is again inelastic. The exterior tough and unyielding matter is for the firmness of the connexion of the several vertebrÆ with each other; the interior softer and elastic matter is for the easy play of the vertebrÆ upon each other; the one for security, the other for pliancy. And the adjustment of the one to the other is such as to combine these properties in a perfect, manner. The quantity of the unyielding substance is not so great as to produce rigidity; the quantity of the elastic substance is not so great as to occasion insecurity. The firm union of its solid matter renders the entire column strong; the aggregate elasticity of its softer substance renders it springy.

Fig. XLVII.
1. One of the Lumbar vertebrÆ. 2. Intervertebral substance. 3. A portion of the spinal cord in its canal.

82. The column is not constructed in such a manner as to admit of an equal degree of motion in every part of it. Every thing is contrived to give to that portion which belongs to the neck freedom of motion, and, on the contrary, to render that portion which belongs to the back comparatively fixed. In the neck the mechanism of the articulating processes is such as to admit of an equal degree of sliding motion forwards, backwards, and from side to side, together with a turning motion of one bone upon another; at the same time, the intervertebral substance between the several vertebrÆ is thick. In consequence of this mechanism, we can touch the breast with the chin, the back with the hind head, and the shoulders with the ear, while we can make the head describe more than a semicircle. But, in the back, the articulating processes are so connected as to prevent the possibility of any motion, either forwards or backwards, or any turning of one vertebra upon another, while the intervertebral substance is comparatively thin (fig. XLV. 2, 4). That portion of the column which belongs to the back is intended to afford a fixed support for the ribs, a support which is indispensable to their action in the function of respiration. In the loins, the articulating processes are so connected as to admit of a considerable degree of motion in the horizontal direction, and from side to side, and the intervertebral substance here progressively increases in thickness to the point at which the upper portion of the column is united to the sacrum (fig. XLV. 2, 4), where the degree of motion is extensive.

83. The canal for the spinal cord, formed partly by the concavity in the posterior surface of the vertebra, and partly by the arch that springs from it (fig. XLVI. 6.), is lined by a continuation of the dense and strong membrane that constitutes the internal periosteum of the cranium, the dura mater (fig. XLVIII. c), which, passing out of the opening in the occipital bone, called the foramen magnum (figs. XXXIX. 7, and XLIX. 3), affords a smooth covering to the canal throughout its whole extent.

Fig. XLVIII
a. The scalp, turned down.
b. The cut edge of the bones of the skull.
c. The external strong membrane of the brain (Dura Mater)
suspended by a hook.
d. The left hemisphere of the brain, showing its convolutions.
e. The superior edge of the right hemisphere.
f. The fissure between the two hemispheres.

Fig. XLIX
1. Hemispheres of the brain proper, or cerebrum;
2. hemispheres of the smaller brain, or cerebellum; 3. spinal
cord continuous with the brain, and the spinal nerves proceeding
from it on each side.

84. The spinal cord itself, continuous with the substance of the brain, passes also out of the cranium through the foramen magnum into the spinal canal (fig. XLIX. 3), enveloped in the delicate membranes that cover it, and surrounded by the aqueous fluid contained between those membranes. The size of the spinal canal, accurately adapted to that of the spinal cord, which it lodges and protects, is of considerable size, and of a triangular shape in its cervical portion (fig. XLIX. 3), smaller and rounded in its dorsal portion (fig. XLIX. 3), and again large and triangular in its lumbar portion (fig. XLIX. 3).

85. The spinal column performs several different, and apparently incompatible, offices.

First, it affords a support and buttress to other bones. It sustains the head (fig. XXXIV. 1); it is a buttress to the ribs (fig. XLVI. 7); through the sternum and ribs it is also a buttress to the superior, and through the pelvis, to the lower, extremities (fig. XXXIV. 2, 3, 4).

Secondly, it affords a support to powerful muscles, partly to those that maintain the trunk of the body in the erect posture against the force of gravitation, and partly to those that act upon the superior and inferior extremities in the varied, energetic, and sometimes long-continued movements they execute.

Thirdly, it forms one of the boundaries of the great cavities that contain the chief organs of the organic life. To the support and protection of those organs it is specially adapted; hence the surface in immediate contact with them is even and smooth; hence its different curvatures, convexities, and concavities, have all reference to their accommodation; hence in the neck it is convex (fig. XLV. 2), in order to afford a firm support to the esophagus, the wind-pipe, the aorta, and the great trunks of the venous system (fig. LX. 3, 4); in the back it is concave, in order to enlarge the space for the dilatation of the lung in the act of inspiration (figs. XLV. 3, and LX. 5); in the loins it is convex, in order to sustain and fix the loose and floating viscera of the abdomen (figs. XLV. 4, and LX. 6, 7, 8, 9); in the pelvis it is concave, in order to enlarge the space for lodging the numerous delicate and highly-important organs contained in that cavity (fig. XLV. 5).

Fourthly, it forms the osseous walls of a canal (figs. XLVI. 6, and XLVII. 3) for the lodgment and protection of the soft and tender substance of the spinal cord, one of the great central masses of the nervous system, the seat of the animal life (fig. XLIX. 3).

Fifthly, it affords in its osseous walls secure apertures for the passage of the spinal nerves (figs. XLVI. 7, and XLIX. 3), by which impressions are transmitted from the organs to the spinal cord and brain, in the function of sensation; and from the spinal cord and brain to the organs in the function of volition.

86. For the due performance of these offices, it is indispensable that it should be firm, rigid, strong, and yet to a certain extent readily flexible in every direction. By what mechanism is it endowed with these apparently incompatible properties?

87. By means of the ring of compact bone, which forms so large a part of its body (fig. XLVI. 1) it is rendered firm, rigid, and strong. By means of its numerous separate pieces, exactly adjusted to each other, and dove-tailed into one another, an increase of strength is gained, such as it would not have been possible to communicate to a single solid piece. By the same mechanism, some degree of flexibility is also obtained; each separate bone yielding to some extent, which, though slight in a single bone, becomes considerable in the twenty-four.

88. But the flexibility required is much greater than could be obtained by this expedient alone. A rigid and immoveable pile of bones, in the position of the spinal column, on which all the other parts of the body rest, and to which they are directly or indirectly attached, would necessarily have rendered all its movements stiff and mechanical; and every movement of every kind impossible, but in a given direction. That the movements of the body may be easy, free, and varied; that it may be possible to bring into play new and complex combinations of motion at any instant, with the rapidity of the changes of thought, at the command of the impulses of feeling, it is indispensable that the spinal column be flexible in every direction, forwards, backwards, and at the sides: it is equally indispensable that it be thus capable of yielding, without injuring the spinal cord; without injuring the spinal nerves; without injuring the thoracic and abdominal viscera; and without injuring the muscles of the trunk and extremities. The degree in which it possesses this power of flexibility, and the extent to which, by the cultivation of it, it is sometimes actually brought, is exemplified in the positions and contortions of the posture-master and the tumbler. It is acquired by means of the intervertebral substance, the compressible and elastic matter interposed between the several vertebrÆ. So compressible is this substance, that the human body is half an inch shorter in the evening than in the morning, having lost by the exertions of the day so much of its stature; yet, so elastic is this matter, that the stature lost during the day is regained by the repose of the night. The weight of the body pressing in all directions upon the spinal column; muscles, bones, cartilages, ligaments, membranes, with all their vessels and all the fluids contained in them; the weight of all these component parts of the head, trunk, and extremities, pressing, without the cessation of an instant, during all the hours of vigilance, upon the intervertebral substance, compresses it; but this weight, being taken off during the night, by the recumbent posture of the body, the intervertebral substance, in consequence of its elasticity, regains its original bulk, and of course the spinal column its original length.

89. But the flexibility acquired through the combined properties of compressibility and elasticity is exceedingly increased by the action of the pulpy and inelastic matter in the centre of the intervertebral substance; this matter serving as a pivot to the vertebrÆ, facilitating their motion on each other. Its effect has been compared to that of a bladder partly filled with water, placed between two trenchers; in this case, the approximation of the circumference of the two trenchers on one side, would instantly displace a portion of the water on that side, which would occupy the increasing space on the other, with the effect of facilitating the change, in every possible direction, of the position of the two trenchers in relation to each other. To this effect, however, it is indispensable that the matter immediately around this central pivot should be, not like itself, rigid and unyielding, but compressible and elastic. It is an interesting fact, that since this illustration was suggested, it has been discovered that this very arrangement is actually adopted in the animal body. In certain animals, in the very centre of their intervertebral substance, there has been actually found a bag of water, with a substance immediately surrounding the bag, so exceedingly elastic, that when the bag is cut, the fluid contained in it is projected to the height of several feet in a perpendicular stream.

90. But besides securing freedom and extent of motion, the intervertebral substance serves still another purpose, which well deserves attention.

Firmness and strength are indispensable to the fundamental offices performed by the column; and to endow it with these properties, we have seen that the external concentric layers of the intervertebral substance are exceedingly tough and that they are attached to the bodies of the vertebrÆ, which are composed of dense and compact bone. But than dense and compact bone, nothing can be conceived better calculated to receive and transmit a shock or jar on the application of any degree of force to the column. Yet such force must necessarily be applied to it in every direction, from many points of the body, during almost every moment of the day; and did it actually produce a corresponding shock, the consequence would be fatal: the spinal cord and brain would be inevitably killed; for the death of these tender and delicate substances may be produced by a violent jar, although not a particle of the substances themselves be touched. A blow on the head may destroy life instantaneously, by what is termed concussion; that is, by the communication of a shock to the brain through the bones of the cranium. The brain is killed; but on careful examination of the cerebral substance after death, not the slightest morbid appearance can be detected: death is occasioned merely by the jar. A special provision is made against this evil, in the structure of the bones of the cranium, by the interposition between its two compact plates of the spongy substance called diploË (fig. XLIII. 2); and this is sufficient to prevent mischief in ordinary cases. A great degree of violence applied directly to the head is not common: when it occurs it is accidental: thousands of people pass through life without ever having suffered from it on a single occasion: but every hour, in the ordinary movements of the body, and much more in the violent movements which it occasionally makes, a degree of force is applied to the spinal column, and through it transmitted to the head, such as, did it produce a proportionate shock, would inevitably and instantly destroy both spinal cord and brain. The evil is obviated partly by the elastic, and partly by the non elastic properties of the matter interposed between the several layers of compact bone. By means of the elastic property of this matter, the head rides upon the summit of the column as upon a pliant spring, while the canal of the spinal cord remains secure and uninvaded. By means of the soft and pulpy portion of this matter, the vibrations excited in the compact bone are absorbed point by point as they are produced: as many layers of this soft and pulpy substance, so many points of absorption of the tremors excited in the compact bone; so many barriers against the possibility of the transmission of a shock to the delicate nervous substance.

91. Alike admirable is the mechanism by which the separate pieces of the column are joined together. If but one of the bones were to slip off its corresponding bone, or to be displaced in any degree, incurable paralysis, followed ultimately by death, or instantaneous death, would happen; for pressure on the spinal cord in a certain part of its course is incompatible with the power of voluntary motion, and with the continuance of life for any protracted term; and in another part of its course, with the maintenance of life beyond a few moments. To prevent such consequences, so great is the strength, so perfect the attachment, so unconquerable the resistance of that portion of the intervertebral substance which surrounds the edge of the bodies of the vertebrÆ, that it will allow the bone itself to give way rather than yield. Yet such is the importance of security to this portion of the frame, that it is not trusted to one expedient alone, adequate as that might seem. Besides the intervertebral substance, there is another distinct provision for the articulation of the bodies of the vertebrÆ. Commencing at the second cervical vertebra, in its fore part, and extending the whole length of the column to the sacrum, is a powerful ligament, composed of numerous distinct longitudinal fibres (fig. L.), which are particularly expanded over the intervals between the bones occupied by the intervertebral substance (figs. L. 1, and LI. 2, 2). This ligament is termed the common anterior vertebral, beneath which, if it be raised from the intervertebral substance, may be seen small decussating fibres, passing from the lower edge of the vertebra above, to the upper edge of the vertebra below (fig. L. 3), from which circumstance these fibres are termed crucial.

Fig. L.
1. Common anterior ligament; 2. intervertebral substance. The anterior ligament is removed to exhibit (3.) the crucial fibres passing over it.
Fig. LI.
1. Portion of the occipital bone; 2. common anterior ligament.

92. Corresponding with the ligament on the anterior, is another on the posterior part of the spine (fig. LII. 1), which takes its origin from the foramen magnum (fig. LII. 1); descends from thence, within the vertebral canal, on the posterior surface of the bodies of the vertebra (fig. LII. 1), and extends to the sacrum. This ligament is termed the common posterior vertebral, which, besides adding to the strength of the union of the bodies of the vertebrÆ, prevents the column itself from being bent too much forward.

Fig. LII.
1. Posterior vertebral ligament.

93. Moreover, the bony arches of the vertebrÆ (fig. LIII. 1) are connected by means of a substance partly ligamentous, and partly cartilaginous (fig. LIII. 2), which, while it is extremely elastic, is capable of resisting an extraordinary degree of force.

Fig. LIII.
1. Arches of the vertebrÆ seen from within;
2. ligaments connecting them.

94. And in the last place, the articular processes form so many distinct joints, each being furnished with all the apparatus of a moveable joint, and thus possessing the ordinary provision for the articulation of bones, in addition to the whole of the foregoing securities.

95. "In the most extensive motion of which the spinal column is capable, that of flexion, the common anterior ligament is relaxed; the fore part of the intervertebral substance is compressed, and its back part stretched; while the common posterior ligament is in a state of extension. In the extension of the column the state of the ligaments is reversed; those which were extended being in their turn relaxed, while the common anterior vertebral is now put upon the stretch. In the lateral inclination of the column, the intervertebral substance is compressed on that side to which the body is bent. In the rotatory motion of the column, which is very limited in all the vertebrÆ, but more particularly in the dorsal, in consequence of their attachment to the ribs, the intervertebral substance is contorted, as are likewise all the ligaments. All the motions of the column are capable of being aided to a great extent by the motion of the pelvis upon the thighs."

96. "The number and breadth of the attachments of these bones," says an accomplished anatomist and surgeon,[4] "their firm union by ligament, the strength of their muscles, the very inconsiderable degree of motion which exists between any two of them, and lastly, the obliquity of their articular processes, especially in the dorsal and lumbar vertebrÆ, render dislocation of them, at least in those regions, impossible without fracture; and I much doubt whether dislocation even of the cervical vertebrÆ ever occurs without fracture, either through their bodies or their articular processes. The effects of each of these accidents would produce precisely the same injury to the spinal marrow, and symptoms of greater or less importance, according to the part of the spinal column that is injured. Death is the immediate consequence if the injury be above the third cervical vertebra, the necessary paralysis of the parts to which the phrenic and intercostal nerves are distributed causing respiration instantly to cease. If the injury be sustained below the fourth cervical vertebra, the diaphragm is still capable of action, and dissolution is protracted. The symptoms, in fact, are less violent in proportion as the injury to the spinal marrow is further removed from the brain; but death is the inevitable consequence, and that in every case at no very distant period."

97. So the object of the construction of the spinal column being to combine extent and freedom of motion with strength, and it being necessary to the accomplishment of this object to build up the column of separate pieces of bone, the connecting substances by which the different bones are united are constituted and disposed in such a manner as to prove absolutely stronger than the bones themselves. Such is the structure of this important portion of the human body considered as a piece of mere mechanism; but our conception of its beauty and perfection would be most inadequate if we did not bear in mind, that while the spinal column performs offices so varied and apparently so incompatible, it forms an integrant portion of a living machine: it is itself alive: every instant, blood-vessels, absorbents and nerves, are nourishing, removing, renewing, and animating every part and particle of it.

98. The anterior boundary of the thorax is formed by the bone called the sternum, or the breast-bone, which is broad and thick at its upper, and thin and elongated at its lower extremity (figs. XLV. 6, and LIV.), where it gives attachment to a cartilaginous appendix, which being pointed and somewhat like a broadsword, is called the ensiform cartilage.

99. Its position is oblique, being near the vertebral column at the top, and distant from it at the bottom (fig. XLV. 6). Its margins are thick, and marked by seven depressions, for the reception of the cartilages of the seven true ribs (fig. LIV). Its anterior surface is immediately subjacent to the skin, and gives attachment to powerful muscles, which act on the superior extremities: its posterior surface is slightly hollowed in order to enlarge the cavity of the thorax (fig. LV.).

Fig. LV.
Posterior view of the sternum.

100. The thorax is bounded at the sides by the ribs, which extend like so many arches between the spinal column and the sternum (fig. XLV. 7, 8). They are in number twenty-four, twelve on each side, of which the seven upper are united to the sternum by cartilage, and are called true ribs (fig. XLV. 7); the cartilages of the remaining five are united with each other and are not attached to the sternum; these are called false ribs (fig. XLV. 8): all of them are connected behind to the spinal column (fig. XXXV.).

101. The ribs successively and considerably increase in length as far as the seventh, by which the cavity they encompass is enlarged; from the seventh they successively diminish in length, and the capacity of the corresponding part of the cavity is lessened. The direction of the ribs from above downwards is oblique (fig. XLV. 7, 8). Their external or anterior surface is convex (fig. XLV. 7, 8); their internal or posterior surface is concave: by the first their strength is increased; by the second the general cavity of the thorax is enlarged (fig. XLV. 7, 8). Their upper margin is smooth and rounded, and gives attachment to a double layer of muscles, called the intercostal, placed in the intervals that separate the ribs from each other (fig. LIX.). Along the lower margin is excavated a deep groove, for the lodgment and protection of the intercostal vessels.

102. The ribs are connected with the spinal column chiefly by what is termed the anterior ligament (fig. LVI. 1), which is attached to the head of the rib (fig. LVI.), and which, dividing into three portions (fig. LVI. 1), firmly unites every rib to two of the vertebrÆ, and to the intervertebral substance (fig. LVI. 1). This articulation is fortified by a second ligament (fig. LVI. 2), also attached to a head of the rib, termed the interarticular (fig. LVI. 2), and by three others, one of which is attached on the fore part, and the two others in the back part, to the neck of the rib (fig. LVII. 1).

Fig. LVI.
Ligaments connecting the ribs to the spinal column.
1. anterior ligaments; 2. interarticular ligament; 3. ligaments
of the necks of the ribs.

The cartilages of the seven superior ribs are attached to the sternum by a double layer of ligamentous fibres, termed the anterior and the posterior ligaments of the sternum (fig. LVIII.). So strong are the bands which thus attach the ribs to the spinal column and the sternum, that the ribs cannot be dislocated without fracture. "Such at least is the case in experiments upon the dead body, where, though the rib be subjected to the application of force by means of an instrument best calculated to detach its head from the articulation, yet it is always broken."

Fig. LVII.
1, &c. Ligaments connecting the ribs to the vertebrÆ behind.

While thus firmly attached to their points of support, the ligaments, which fix them, are so disposed as to render the ribs capable of being readily moved upwards and downwards: upwards in inspiration; downwards in expiration; and it is by this alternate action that they enlarge and diminish the cavity of the thorax in the function of respiration.

Fig. LVIII.
Ligaments joining the cartilages of the ribs to the sternum.

103. Such are the boundaries of the cavity of the thorax as far as its walls are solid. The interspaces between these solid portions at the sides are filled up by muscles, principally by those termed the intercostal (fig. LIX.); below, the boundary is formed by the diaphragm (fig. LXI. 2); while above, as has been already stated (69), the cavity is so contracted as only to leave an opening for the passage of certain parts to and from the chest.

Fig. LIX.
A view of the muscles called Intercostals, filling up the
spaces between the ribs.

104. The inner surface of the walls of the thorax, in its whole extent, is lined by a serous membrane, exceedingly thin and delicate, but still firm, called the pleura. The same membrane is reflected over the organs of respiration contained in the cavity, so as to give them an external coat. The membrane itself is every where continuous, and every where the same, whether it line the containing or the contained parts; but it receives a different name as it covers the one or the other: that portion of it which lines the walls of the cavity being called the costal pleura (fig. LXI. a), while that which covers the organs contained in the cavity is termed the pulmonary pleura (fig. LX. 5, 1).

105. A fold of each pleura passes directly across the central part of the cavity of the thorax; extending from the spinal column to the sternum, and dividing the general cavity into two. This portion of the pleura is called the mediastinum, from its situation in the centre of the thorax, and it so completely divides the thoracic cavity into two, that the organs on one side of the chest have no communication with those of the other; so that there may be extensive disease in one cavity (for example, a large accumulation of water,) while the other may be perfectly sound.

106. The main organs contained in the cavity of the thorax are the lungs with their air tube; the heart with its great vessels; and the tube passing from the mouth to the stomach (fig. LX.).

107. The two lungs occupy the sides of the chest (fig. LX. 5). They are completely separated from each other by the membranous partition just described, the mediastinum. Between the two folds of the mediastinum, namely, in the middle of the chest, but inclining somewhat to the left side, is placed the heart, enveloped in another serous membrane, the pericardium (fig. LX. 2, 1).

108. The lungs are moulded to the cavities they fill; whence their figure is conical, the base of the cone being downwards, resting on the diaphragm (fig. LX. 5, b); and the apex upwards, towards the neck (fig. LX. 5).

109. That surface of each lung which corresponds to the walls of the chest is convex in its whole extent (fig. LX. 5); on the contrary, that surface which corresponds to the mediastinum is flattened (fig. LX. 5). The basis of the lung is concave, adapted to the convexity of the diaphragm on which it rests (fig. LX. 5).

110. The air-vessel of the lungs, termed the bronchus, together with the blood-vessels and nerves, enter the organ at its flattened side, not exactly in the middle, but rather towards the upper and back part. This portion is termed the root of the lung.

111. The lungs are attached to the neck by the trachea (fig. LX. 4), the continuation of which forms the bronchus; to the spinal column by the pleura, and to the heart by the pulmonary vessels (fig. LX. 3, d): their remaining portion is free and unattached.

112. In the living body, the lungs on each side completely fill the cavity of the chest, following passively the movements of its walls, and accurately adapting themselves to its size, whether its capacity enlarge in inspiration, or diminish in expiration, so that the external surface of the lung (the pulmonary pleura) is always in immediate contact with the lining membrane of the walls of the cavity (the costal pleura); consequently, during life, there is no cavity, the chest being always completely full.

Fig. LX.
a. The cut edges of the ribs, forming the lateral boundaries of the
cavity of the thorax.
b. The diaphragm, forming the inferior boundary of the thorax, and
the division between the thorax and the abdomen.
c. The cut edges of the abdominal muscles, turned aside, exposing
the general cavity of the abdomen.
1. The cut edge of the pericardium
turned aside.
2. The heart.
3. The great vessels in immediate
connexion with the
heart.
4. The trachea, or wind-pipe.
5. The lungs.
6. The liver.
7. The stomach.
8. The large intestine.
9. The small intestines.
10. The urinary bladder.

113. The anterior surface of the pericardium, the bag which envelopes the heart, lies immediately behind the sternum, and the cartilages of the second, third, fourth, and fifth ribs, covered at its sides by the pleura, and firmly attached below to the diaphragm (fig. LX. 1).

114. Surrounded by its pericardium, within the mediastinum, the heart is placed nearly in the centre of the chest, but its direction is somewhat oblique, its apex being directly opposite to the interval between the fifth and sixth ribs on the left side (fig. LX. 2); while its basis is directed upwards, backwards, and towards the right (fig. LX. 2). That portion of its surface which is presented to view on opening the pericardium is convex (fig. LX. 2); but its opposite surface, namely, that which rests upon the part of the pericardium which is attached to the diaphragm, is flattened (fig. LX. 1). It is fixed in its situation partly by the pericardium and partly by the great vessels that go to and from it. But under the different states of expiration and inspiration, it accompanies, in some degree, the movements of the diaphragm; and in the varied postures of the body, the heart deviates to a certain extent from the exact position here described.

115. The second division of the trunk, the abdomen, is bounded above by the diaphragm (fig. LXI. 2), below by the pelvis (fig. LXI. 3), behind and at the sides by the vertebrÆ and muscles of the loins (fig. LXIII.), and before by the abdominal muscles (fig. LXIII. 9).

116. The organ which forms the superior boundary of the abdomen, the diaphragm (midriff), is a circular muscle, placed transversely across the trunk, nearly at its centre (fig. LXI. 2). It forms a vaulted partition between the thorax and the abdomen (fig. LXI. 2). All around its border it is fleshy (fig. LXI. 2); towards its centre it is tendinous (fig. LXI. 2); the surface towards the abdomen is concave (fig. LXI. 2); that towards the thorax convex (fig. LXI. 2); while its middle tendinous portion ascends into the thorax as high as the fourth rib (fig. LXI. 2).

Fig. LXI.
View of the diaphragm. 1. Cavity of the thorax;
2. diaphragm separating the cavity of the thorax from that of
the abdomen; 3. cavity of the pelvis.

117. The diaphragm is perforated by several apertures, for the transmission of tubes and vessels, which pass reciprocally between the thorax and abdomen (fig. LXII.).

1. A separate aperture is formed to afford an exit from the thorax of the aorta, the common source of the arteries (fig. LXII. 2), and an entrance into the thorax of the thoracic duct, the tube that bears the digested aliment to the heart. 2. A little to the left of the former, there is another aperture, through which passes the esophagus or gullet (fig. LXII. 3), the tube that conveys the food from the mouth to the stomach. 3. On the right side, in the tendinous portion of the diaphragm, very carefully constructed, is a third aperture for the passage of the vena cava (fig. LXII. 4), the great vessel that returns the blood to the heart from the lower parts of the body.

Fig. LXII.
View of the diaphragm with the tubes that pass through
it. 1. Arch of the diaphragm; 2. the trunk of the aorta
passing from the chest into the abdomen; 3. the esophagus
passing from the chest through the diaphragm to the stomach;
4. the vena cava, the great vein that returns the
blood to the heart from the lower parts of the body, passing
from the abdomen, into the chest, in its way to the right
side of the heart; 5. 6. muscles that arise in the interior of
the trunk and that act upon the thigh; 5. the muscle called
psoas; 6. the muscle called iliacus.

118. The partition formed by the diaphragm between the thorax and abdomen, though complete, is moveable; for as the diaphragm descends in inspiration and ascends in expiration, it proportionally enlarges or diminishes the cavities between which it is placed; consequently, the actual magnitude of these cavities varies every moment, and the size of the one is always in the inverse ratio of that of the other.

119. Between the abdomen and the pelvis there is no separation; one cavity is directly continuous with the other (fig. LXI. 3); but along the inner surface of the expanded bones, which form a part of the lateral boundary of the abdomen, there is a prominent line, termed the brim of the pelvis (fig. XLV. 15), marking the point at which the abdomen is supposed to terminate and the pelvis to commence.

120. Behind and at the sides the walls of the abdomen are completed partly by the lumbar portion of the spinal column and partly by the lumbar muscles (fig. XLV. 4), and before by the abdominal muscles (fig. LXIII. 9).

121. The inner surface of the walls of the abdomen is lined throughout by a serous membrane, termed the peritoneum (fig. LXIII.). From the walls of the abdomen, the peritoneum is reflected upon the organs contained in the cavity, and is continued over them so as to form their external coat. The peritoneum also descends between the several organs, connecting them together, and holding them firmly in their situation; and it likewise forms numerous folds, in which are embedded the vessels and nerves that supply the organs. It secretes a serous fluid, by which its own surface and that of the organs it covers is rendered moist, polished, and glistening, and by means of which the organs glide smoothly over it, and over one another in the various movements of the body, and are in constant contact without growing together. In structure, distribution, and function, the peritoneum is thus perfectly analogous to the pleura.

122. Like the thorax, the abdomen is always completely full. When the diaphragm is in action, it contracts. When the diaphragm is in the state of contraction, the abdominal and lumbar muscles are in the state of relaxation. By the contraction of the diaphragm, the organs contained in the abdomen are pushed downwards, and the anterior and lateral walls of the cavity being at this moment in a state of relaxation, they readily yield, and, consequently, the viscera are protruded forwards and at the sides. But the abdominal and lumbar muscles in their turn contract, the diaphragm relaxing; and, consequently, the viscera, forced from the front and sides of the abdomen, are pushed upwards, together with the diaphragm, into the cavity of the thorax. A firm and uniform pressure is thus at all times maintained upon the whole contents of the abdomen: there is an exact adaptation of the containing to the contained parts, and of one organ to another. No space intervenes either between the walls of the abdomen and the organs they enclose, or between one organ and another: so that the term cavity does not denote a void or empty space, but merely the extent of the boundary within which the viscera are contained.

123. The contents of the abdomen consist of the organs which belong to the apparatus of digestion, and of those which belong to the apparatus of excretion.

124. The organs which belong to the apparatus of digestion are—1. The stomach (fig. LXIII. 2) 2. The duodenum (fig. LXIII. 4). 3. The jejunum (fig. LXIII. 5). 4. The ilium (fig. LXIII. 5). The three last organs are called the small intestines, and their office is partly to carry on the digestion of the aliment commenced in the stomach, and partly to afford an extended surface for the absorption of the nutriment as it is prepared from the aliment. 5. The pancreas (fig. LXIV. 5). 6. The liver (fig. LXIV. 2). 7. The spleen (fig. LXIV. 4). The three last organs truly belong to the apparatus of digestion, and their office is to co-operate with the stomach and the small intestines in the conversion of the aliment into nutriment.

Fig. LXIII.
1. Esophagus; 2. stomach; 3. liver raised, showing its
under surface; 4. duodenum; 5. small intestines; 6. cÆcum; 7. colon; 8. urinary bladder; 9. gall bladder;
10. abdominal muscles divided and reflected.

125. The organs which belong to the apparatus of excretion are—1. The large intestines consisting of the cÆcum (fig. LXIII. 6). 2. The colon (fig. LXIII. 7). 3. The rectum (fig. LXIV. 10). It is the office of these organs, which are called the large intestines, to carry out of the system that portion of the alimentary mass which is not converted into nourishment. 4. The kidneys (fig. LXIV. 6), the organs which separate in the form of the urine an excrementitious matter from the blood, in order that it may be conveyed out of the system.

Fig. LXIV.
General view of the viscera of the abdomen. 1. Stomach
raised; 2. under surface of liver; 3. gall bladder; 4. spleen;
5. pancreas; 6. kidneys; 7. ureters; 8. urinary bladder;
9. portion of the intestine called duodenum; 10. portion of
the intestine called rectum; 11. the aorta.

126. The last division of the trunk, called the pelvis (fig. LXI. 3), consists of a circle of large and firm bones, interposed between the lower portion of the trunk and the inferior extremities (fig. XLV.). The bones that compose the circle, distinct in the child, are firmly united in the adult into a single piece; but as the original separation between each remains manifest, they are always described as separate bones. They are the sacrum (fig. XLV. 5), the coccyx (fig. XXXV.), the ilium (fig. XLV. 11), the ischium (fig. XLV. 12), and the pubis (fig. XLV. 13).

127. The sacrum, placed like a wedge between the moveable portion of the spinal column and the lower extremities, forms the posterior boundary of the pelvis. The figure of this bone is triangular (fig. XLV. 5); its anterior surface is concave and smooth, for enlarging the cavity of the pelvis and sustaining the organs contained in it (fig. XLV. 5); its posterior surface is convex, irregular, and rough (fig. XXXV.), giving origin to the great muscles that form the contour of the hip, and to the strong muscles of the back and loins that raise the spine and maintain the trunk of the body erect.

128. The base or upper part of the sacrum receives the last vertebra of the loins on a large and broad surface (fig. XLV. 4), forming a moveable joint; and the degree of motion at this point is greater than it is at the higher points of the spinal column. Firmly united at its sides with the haunch bones, it admits there of no degree of motion.

129. The coccyx, so named from its resemblance to the beak of the cuckoo, when elongated by a succession of additional bones, forms the tail in quadrupeds; but in man it is turned inwards to support the parts contained in the pelvis, and to contract the lower opening of the cavity. By means of a layer of cartilage, the medium by which this bone is connected with the sacrum, it forms a moveable articulation, continuing moveable in men until the age of twenty-five, and in women until the age of forty-five; continuing moveable in women thus long, in order that by yielding to the force which tends to push it backwards during the period of labour, it may enlarge the lower aperture of the pelvis, and so facilitate the process of parturition and diminish its suffering.

130. The lateral boundaries of the pelvis are formed by the ilium, the haunch bone (fig. XLV. 11), and by the ischium, the hip bone (fig. XLV. 12). The ilium forms the lower part of the abdomen and the upper part of the pelvis (fig. XLV. 11); its broad expanded wing supports the contents of the abdomen, and gives attachment to the muscles that form the anterior portion of its walls (figs. XLV. 11, and LXIII. 9); its external convex surface sustains the powerful muscles that extend the thigh; and along its internal surface is the prominent line which marks the brim of the pelvis (fig. XLV. 15), and which divides this cavity from that of the abdomen.

131. The ischium or hip bone is the lower part of the pelvis (fig. XLV. 12); at its undermost portion is a rounded prominence called the tuberosity (fig. XLV. 12), in its natural condition covered with cartilage, upon which is superimposed a cushion of fat. It is this part on which the body is supported in a sitting posture.

132. The pubis or share bone forms the upper and fore part of the pelvis (fig. XLV. 13), and together with the two former bones, completes the large and deep socket, termed the acetabulum (fig. XLV. 14), into which is received the head of the thigh-bone (fig. XXXIV. 4). The margin of the acetabulum and the greater part of its internal surface is lined with cartilage, so that in its natural condition it is much deeper than it appears to be when the bones alone remain.

133. The lower aperture of the pelvis, which appears large when all the soft parts are removed, is not really large, for in its natural state it is filled up partly by muscles and partly by ligaments, which sustain and protect the pelvic organs, leaving only just space enough for the passage to and from those which have their opening on the external surface.

134. The cavity of the pelvis, together with all the organs contained in it, are lined by a continuation of the membrane that invests the abdomen and its contents.

135. The organs contained in the pelvis are the rectum (fig. LXIV. 9), which is merely the termination of the large intestines, the urinary bladder (fig. LXIV. 8), and the internal part of the apparatus of reproduction.

136. The large and strong bones of the pelvis not only afford lodgment and protection to the tender organs contained in its cavity, but sustain the entire weight of the body, the trunk resting on the sacrum as on a solid basis (fig. XLV. 5), and the lower extremities being supported in the sockets in which the heads of the thigh-bones play, in the varied movements of locomotion (fig. XXXIV. 4).

137. The last division of the body comprehends the superior and the inferior extremities.

138. The superior extremities consist of the shoulder, arm, fore-arm, and hand.

139. The soft parts of the SHOULDER are composed chiefly of muscles; its bones are two, the scapula or the blade bone, and the clavicle or the collar bone (fig. LXV. 2, 4).

Fig. LXV.
1. Sternum; 2. clavicle; 3. ribs; 4. anterior surface of
scapula; 5. coracoid process of scapula; 6. acromion process
of scapula; 7. margin of glenoid cavity of scapula;
8. body of the humerus or bone of the arm; 9. head of the
humerus.

140. The SCAPULA is placed upon the upper and back part of the thorax, and occupies the space from the second to the seventh ribs (fig. LXV. 4)

Fig. LXVI.
1. Posterior surface of scapula; 2. margin of scapula;
3. acromion process; 4. margin of glenoid cavity; 5. clavicle;
6. body of humerus; 7. head of humerus.

Unlike that of any other bone of the body, it is embedded in muscles, without being attached to any bone of the trunk, excepting at a single point. From the bones of the thorax it is separated by a double layer of muscles, on which it is placed as upon a cushion, and over the smooth surface of which it glides. Originally, like the bones of the skull, it consisted of two tables of compact bone, with an intermediate layer of spongy bony substance (diploË); but, by the pressure of the muscles that act upon it, it gradually grows thinner and thinner, until, as age advances, it becomes in some parts quite transparent and as thin as a sheet of paper.

141. The figure of the scapula is that of an irregular triangle (fig. LXVI.). Its anterior surface is concave (fig. LXV. 4), corresponding to the convexity of the ribs (fig. XLV. 7); its posterior surface is very irregular (fig. LXVI. 1), being in some parts concave and in others convex, giving origin especially to two large processes (figs. LXV. 5, and LXVI. 3); one of which is termed the acromion (fig. LXVI. 3), and the other the coracoid process of the scapula (fig. LXV. 5). The margins of the bone, whatever the thinness of some portions of it, are always comparatively thick and strong (fig. LXVI. 2), affording points of origin or of insertion to powerful muscles. At what is called the anterior angle of the bone there is a shallow oval depression covered with cartilage and deepened by a cartilaginous margin, called the glenoid cavity of the scapula (figs. LXV. 7, and LXVI. 4), which receives the head of the humerus or bone of the arm (figs. LXV. 9, and LXVI. 7, 6).

142. The clavicle, the second bone of the shoulder, is a long and slender bone, of the form of an italic , projecting a little forwards towards its middle, so as to give a slight convexity of outline to the top of the chest and the bottom of the neck (fig. LXV. 2). It is attached by one extremity to the sternum (fig. LXV. 2) and by the other to the scapula (fig. LXV. 2), by moveable joints. The nature of an immoveable joint has been explained (63). In the connexion of the bones of the trunk, while the main object is to secure firmness of attachment, some degree of motion is at the same time obtained (81 et seq.): but the mode in which the several bones of the extremities are connected with each other and with the trunk, admits of so great a degree of motion, that these articulations are pre-eminently entitled to the name of moveable joints. The component parts of all moveable joints are bone, cartilage, synovial membrane, and ligament. The great character of a moveable joint is the approximation of two or more bones; yet these bony surfaces are never in actual contact, but are invariably separated from each other by cartilage. The cartilage either covers the entire extent of the articulating surface of the bones, as in the shoulder-joint, where both the head of the humerus and the cavity of the scapula that receives it are enveloped in this substance (fig. LXV. 7. 9), or a portion of it is placed between the articulating surfaces of the bones, as in the joint between the clavicle and sternum (fig. LXVII. a); which, when so placed, is termed an interarticular cartilage (fig. LXVII. a). By its smooth surface cartilage lessens friction; while by its elasticity it facilitates motion and prevents concussion. Slightly organized cartilage is provided with comparatively few blood-vessels and nerves. Had it been vascular and sensible like the skin and the muscle, the force applied in the movements of the joint would have stimulated the blood-vessels to inordinate action, and the sensibility of the nerves would have been the source of constant pain: every motion of every joint would have been productive of suffering, and have laid the foundation of disease. The facility and ease of motion obtained by the smoothness, elasticity, and comparative insensibility of cartilage are still further promoted by the fluid which lubricates it, termed synovia, secreted by a membrane called synovial, which lines the internal surface of the joint, and which bears a close resemblance to the serous (30). Synovia is a viscid fluid of the consistence of albumen (5). It is to the joint what oil is to the wheel, preventing abrasion and facilitating motion; but it is formed by the joint itself, at the moment when needed, and in the quantity required. The motion of the joint stimulates the synovial membrane to secretion, and hence the greater the degree of motion, the larger the quantity of lubricating fluid that is supplied. The several parts of the apparatus of moveable joints are retained in their proper position by ligamentous substance, which, as has been shown (96 and 97), is oftentimes so strong that it is easier to fracture the bone than to tear the ligament, and in every case the kind and extent of motion possessed by the joint are dependent mainly on the form of the articulatory surfaces of the bones and on the disposition of the ligaments.

143. In the joint formed by the clavicle and the sternum (fig. LXVII. a) an interarticular cartilage is placed between the two bones which are united, first by a strong fibrous ligament, which envelops them as in a capsule (fig. LXVII. 1); by a second ligament, which extends from the cartilage of the first rib to the clavicle (fig. LXVII. 4), by which the attachment of the clavicle to the sternum is materially strengthened; and by a third ligament which passes transversely from the head of one clavicle to that of the other (fig. LXVII. 3). The joint thus formed, though so strong and firm that the dislocation of it is exceedingly rare, yet admits of some degree of motion in every direction, upwards, downwards, forwards, and backwards; and this articulation is the sole point by which the scapula is connected with the trunk, and consequently by which the upper extremity can act, or be acted upon, by the rest of the body.

Fig. LXVII.
1. The fibrous capsule of the sternum and clavicle; 2. the
same laid open, showing a, the interarticular cartilage;
3. the ligament connecting the two clavicles; 4. the ligament
joining the clavicle to the first rib; 5. ligaments
passing down in front of the sternum.

144. The scapular extremity of the clavicle (fig. LXVIII. 6) is attached to the processes of the scapula (fig. LXVIII. 4. 3) by several ligaments of great strength (fig. LXVIII. 7, 8, 9). First by very strong fasciculi which pass from the upper surface of the clavicle to the acromion of the scapula (fig. LXVIII. 6); and secondly by two ligaments which unite the clavicle with the coracoid process of the scapula (fig. LXVIII. 8, 9). These ligaments are so powerful that they resist a force capable of fracturing the clavicle; and they need to be thus strong, for the clavicle is a shaft which sustains the scapula, and through the scapula the whole of the upper extremity; and the main object of the joint by which these bones are united, is to afford a firm attachment of the scapula to its point of support.

Fig. LXVIII.
1. The clavicle; 2. the anterior part of the scapula;
3. the coracoid process; 4. the acromion process; 5. the humerus;
6. ligaments binding the scapular end of the clavicle
to the acromion; 7. 8. 9. ligaments passing from one
process of the scapula to the other; 10. the fibrous capsule
of the shoulder-joint.

145. The clavicle serves the following uses: it sustains the upper extremity; it connects the upper extremity with the thorax; it prevents the upper extremity from falling forwards upon the thorax; and it affords a fixed point for steadying the extremity in the performance of its various actions.

146. The glenoid cavity of the scapula (fig. LXV. 7) receives the head of the humerus, the bone of the arm (fig. LXV. 9), and the two bones being united by ligament form the shoulder-joint (fig. LXVIII.). This joint is what is termed a ball and socket joint, the peculiarities of which are two: first, beyond all others this mode of articulation admits of free and extensive motion; in the present case, there is the utmost freedom of motion in every direction, upwards, downwards, backwards, and forwards. In the second place, this mode of articulation admits of the motion of the limb without that of the body, or of the motion of the body without that of the limb. When at rest, the arm may be moved in almost any direction without disturbing the position of any other part of the frame; the manifold advantages of which are obvious. On the other hand, by careful management, very considerable variations in the posture of the body may be effected without the communication of any degree of motion to the limb; an unspeakable advantage when the limb has sustained injury, or is suffering from disease.

147. It does not seem possible to construct a joint of great strength, capable, at the same time, of the degree of motion possessed by the joint of the shoulder. So shallow is the socket of the scapula, and so large the head of the humerus, that it seems as if the slightest movement must dislodge it from its cavity (fig. LXVI. 4. 7). For sustaining heavy weights or resisting a great amount of pressure, applied to it suddenly and in various directions, the arm is obviously unfitted. But this is not its office. The superior extremities are the organs of apprehension—the instruments by which the mind executes the commands of the will. They do not need the strength required by the organs that sustain the weight of the body and that perform the function of locomotion; but they do need freedom and extent of motion: to this strength may be sacrificed, and so it is; yet what can be done to combine strength with mobility is effected. Large and strong processes of bone, proceeding as has been shown (141), from the convex surface of the scapula (figs. LXV. and LXVI.), overhang, and to a considerable extent surround, the head of the arm-bone, especially resisting the force that would dislodge it from its socket and drive it upwards, inwards, and backwards (fig. LXV.), the directions in which force is most commonly applied to it. By these processes of bone the joint is greatly strengthened, especially in those directions. Moreover, a strong ligament, termed the fibrous capsule (fig. LXVIII. 10) envelops the joint. This ligament, arising from the neck of the scapula (fig. LXVIII. 10), expands itself in such a manner as completely to surround the head of the humerus (fig. LXVIII. 10); and then again contracts in order to be inserted into the neck of the bone (fig. LXVIII. 10). This ligament is strengthened by the tendons of no less than four muscles which are expanded over it, as well as by the powerful substance termed fascia which is reflected upon it from both the processes and ligaments of the scapula. In addition to all these expedients for fortifying the joint, it receives a further security in the position of the scapula, which is loose and unattached; which slides easily over the ribs upon its cushion of flesh; which thus obtains, by its facility of yielding, some compensation for its want of strength, eluding the force which it cannot resist.

148. The arm consists of numerous and powerful muscles, and of a single bone, the humerus, which belongs to the class of bones termed cylindrical (185).

149. The upper end of the humerus terminates in a circular head (fig. LXV. 9), which is received into the socket of the scapula (fig. LXV. 9. 7) termed, as has been stated (141), its glenoid cavity. The middle portion of the bone, or what is termed its shaft (fig. LXV. 8), diminishes considerably in magnitude, and becomes somewhat rounded (fig. LXV. 8), while its lower end again enlarges, and is spread out into a flattened surface of great extent (fig. LXIX. 1, 3, 2, 4). Of this broad flattened surface, the middle portion is grooved (fig. LXIX. 2): it is covered with cartilage; it forms the articulating surface by which the arm is connected with the fore-arm. On each side of this groove there is a projection of bone or tubercle, termed condyle (fig. LXIX. 3, 4), the inner (fig. LXIX. 3) being much larger than the outer (fig. LXIX. 4). The inner condyle gives origin to the muscles that bend, the outer to those that extend the fore-arm and the fingers (figs. LXXXIV. 1, 2, and LXXXV. 1).

Fig. LXIX.
1. Lower extremity of the humerus; 2. grooved surface;
3. internal condyle; 4. external condyle; 5. the upper part
of the ulna; 6. the head; 7. the neck; 8. the tubercle of
the radius.

150. The muscles that act upon the arm arise from the back (fig. LXXII. 2), the chest (fig. LXXI. 1), the clavicle (fig. LXXI. 1), and the scapula (fig. LXXI. 3); and they move the arm with freedom and power upwards, downwards, forwards, backwards, inwards, and outwards. The chief muscle that raises the arm is the deltoid (fig. LXXI. 3), which arises partly from the clavicle and partly from the scapula (fig. LXXI. 3). It has the appearance of three muscles proceeding in different directions, the different portions being separated by slight fissures (figs. LXXI. 3, and LXXII. 3). The fibres converging unite and form a powerful muscle which covers the joint of the humerus (fig. LXXI. 3). It is implanted by a short and strong tendon into the middle of the humerus (fig. LXXI. 4). Its manifest action is to pull the arm directly upwards. Its action is assisted by the muscles that cover the back of the scapula, which are in like manner inserted into the humerus, and which, at the same time that they elevate the arm, support it when raised.

Fig. LXXI.
View of the muscles on the fore part of the chest that
act upon the arm. 1. The muscle called the great pectoral;
2. the small pectoral; 3. the deltoid; 4. the humerus.

151. The principal muscle that carries the arm downwards is the latissimus dorsi (fig. LXXII. 2), the broadest muscle of the body, which, after having covered all the lower part of the back and loins, terminates in a thin but strong tendon which stretches to the arm, and is implanted into the humerus (fig. LXXII. 2), near the tendon of a muscle immediately to be described,—the great pectoral. When the arm is raised by the deltoid and its assistant muscles, the latissimus dorsi carries it downwards with force, and its powerful action is increased by that of other muscles which arise from the scapula and are inserted into the arm.

152. The principal muscle that carries the arm forwards towards the chest, is the great pectoral (fig. LXXI. 1), which, arising partly from the clavicle (fig. LXXI. 1), partly from the sternum (fig. LXXI. 1), and partly from the cartilages of the second, third, fourth, fifth, and sixth ribs (fig. LXXI. 1), covers the greater part of the breast (fig. LXXI. 1). Its fibres, converging, terminate in a strong tendon, which is inserted near the tendon of the longissimus dorsi into the humerus, about four inches below its head (fig. LXXI. 1). These two muscles form the axilla or armpit, the anterior border of the axilla consisting of the pectoral muscle. Though each of these muscles has its own peculiar office, yet they often act in concert, thereby greatly increasing their power, and the result of their combined action is to carry the arm either directly downwards or to the side of the chest.

Fig. LXXII.
View of the muscles seated on the back part of the trunk
that act upon the shoulder and arm. 1. The muscle called
the trapezius; 2. the latissimus dorsi; 3. the deltoid.

153. Some of the muscles that elevate the arm carry it inwards, and others outwards; the muscles that carry it forwards likewise carry it inwards; while of the muscles that pull it downwards, some direct it forwards and inwards, and others backwards and outwards (151 and 152).

154. It has been already stated that the shoulder-joint is completely surrounded by the muscular fibres or the tendinous expansions of several of these powerful muscles, which have a far greater effect in maintaining the head of the humerus in its socket than the fibrous capsule of the joint; the latter being necessarily loose, in order to allow of the extended and varied motions of the arm, whereas the muscles that encompass the joint adhere closely and firmly to it. Moreover, by virtue of their vital power these muscles act with an efficiency which a mere ligamentous band is incapable of exerting; for they apportion the strength of resistance to the separating force, and react with an energy proportioned to the violence applied.

155. The bones of the fore-arm are two, the ulna and the radius (figs. LXIX. and LXXIII.). The ulna is essentially the bone of the elbow (figs. LXIX. 5, and LXXIII. 3); the radius that of the hand (fig. LXXV.). Supposing the arm to hang by the side of the body, and the palm of the hand to be turned forwards, the ulna, in apposition with the little finger, occupies the inner; and the radius, in apposition with the thumb, occupies the outer part of the fore-arm (fig. XXXIV. 3).

Fig. LXXIII.
1. The internal condyle of the humerus; 2. the external
condyle of the humerus; 3. the olecranon process of the
ulna; 4. the head of the radius.

156. The upper end of the ulna belonging to the elbow is large (figs. LXIX. 5, and LXXIII. 3). It sends backwards the large projection commonly named the elbow or olecranon (fig. LXXII. 3), in the centre of which there is a smooth and somewhat triangular surface (fig. LXXIII. 3) which is always covered by skin of a coarse texture, like that placed over the lower part of the knee-pan, as if nature intended this for a part on which we may occasionally lean and rest. Large at the elbow, the ulna gradually grows smaller and smaller as it descends towards the wrist, where it ends in a small round head (fig. LXXXII. 2), beyond which, on the inner side, or that corresponding to the little finger, it projects downwards a small rounded point, termed the styloid process (fig. LXXXII. 3). As the styloid process and the olecranon, the two extremities of the ulna (figs. LXXIII. 3, and LXXII. 3), are easily and distinctly felt, the length of this bone was primitively used as a measure, called a cubit, which was the ancient name of the bone.

157. The radius, the second bone of the fore-arm, placed along its outer part next the thumb, is small at its upper end (figs. LXIX. 6, and LXXIII. 4); but its body is larger than that of the ulna; while its lower end, next the wrist to which it properly belongs, is very bulky (fig. LXXXII. 1). Its upper end is formed into a small circular head, which is united by distinct joints both to the humerus and to the ulna (fig. LXIX. 6). The top of its rounded head is excavated into a shallow cup (figs. LXIX. 6, and LXXIII. 4) which receives a corresponding convexity of the humerus (fig. LXIX. 2), and its lower extremity is excavated into an oblong cavity, which receives two of the bones of the wrist (fig. LXXXIII. 1. 4).

158. The joint of the elbow is composed above of the condyles of the humerus (fig. LXIX. 3. 2), and below by the heads of the ulna and radius (fig. LXIX. 5. 6).

159. The upper surface of the ulna is so accurately adapted to the lower surface of the humerus that the one seems to be moulded on the other (figs. LXIX. 5, and LXXIII. 3), and the form of these corresponding surfaces, which are everywhere covered with cartilage, is such as to admit of free motion backwards and forwards, that is, of extension and flexion; but to prevent any degree of motion in any other direction. The joint is therefore a hinge-joint, of which the two motions of flexion and extension are the proper motions. This hinge is formed on the part of the humerus by a grooved surface, with lateral projections (fig. LXIX. 2, 3, 4), and on the part of the ulna by a middle projection with lateral depressions (fig. LXIX. 5): the middle projection of the ulna turning readily on the grooved surface of the humerus (fig. LXIX. 2).

160. The bones are held in their proper situation, first, by a ligament on the fore part of the arm, called the anterior (fig. LXXIV. 6), which arises from the lower extremity of the humerus, and is inserted into the upper part of the ulna and the coronary ligament of the radius (fig. LXXIV. 6. 8); secondly, by another ligament on the back part of the arm, called the posterior ligament (fig.LXXV. 8), placed in the cavity of the humerus that receives the olecranon of the ulna (fig. LXXV. 8); and thirdly, by two other ligaments at the sides of the ulna (fig. LXXV. 6, 7). The ulna and radius are united, first, by a ligament called the coronary, which, arising from the ulna, passes completely around the head of the radius (fig. LXXVI. 3), and the attachment of which, while sufficiently close to prevent the separation of the two bones, is yet not adherent to the radius, for a reason immediately to be assigned; secondly, by another ligament which passes in an oblique direction from one bone to the other (fig. LXXVI. 4); and thirdly, by a dense and broad ligament, termed the interosseous (figs. LXXIV. 10, and LXXVI. 5), which fills up the space between the two bones nearly in their whole extent. This ligament serves other offices besides that of forming a bond of union, affording, more especially, a greater extent of surface for the attachment of muscles, and separating the muscles on the anterior from those on the posterior part of the limb.

161. At their inferior extremities the ulna and radius are united partly by the interosseous ligament (fig. LXXVII. 1) and partly by ligamentous fibres which pass transversely from one bone to the other (fig. LXXVII. 2) on the anterior and the posterior surface of the fore-arm.

Fig. LXXVII.
1. Interosseous ligament; 2. transverse fibres passing
between the radius and ulna, and uniting the two bones;
3. 4. 5. posterior and lateral ligaments of the wrist joint;
6. ligaments uniting the bones of the wrist with one another;
7. 8. ligaments which attach the metacarpal to the
carpal bones; 9. transverse ligaments for the attachment
of the phalanges of the fingers; 10. lateral ligaments for
the attachment of the phalanges of the fingers 11. ligaments
of the thumb.

162. The lower extremity of the radius is also united to the wrist; and the hand being attached to the wrist, the junction of the hand and the fore-arm is effected by the articulation of the wrist with the radius (fig. LXXVII.). The ligaments which connect the bones of the wrist with the radius are bands of exceeding strength (fig. LXXVII. 3).

163. The muscles that act upon the fore-arm are placed upon the arm (fig. LXXVIII.). The joint of the elbow being a hinge-joint, the fore-arm can admit only of two motions, namely, flexion and extension. The muscles by which these motions are effected are four, two for each; the two flexors being placed on the fore part (fig. LXXVIII. 2. 4), and the two extensors on the back part of the arm (fig. LXXIX. 5).

164. The two flexor muscles of the fore-arm are termed the biceps and the brachialis (fig. LXXVIII. 2, 4). The biceps is so called because it has two distinct heads or points of origin (fig. LXXVIII. 2), both of which arise from the scapula (fig. LXXVIII. 2). About a third part down the humerus the two heads meet, unite and form a bulky muscle (fig. LXXVIII. 2), which, when it contracts, may be felt like a firm ball on the fore part of the arm, the upper part of the ball marking the point of union of the two heads (fig. LXXVIII. 2). The muscle gradually becoming smaller, at length terminates in a rounded tendon (fig. LXXVIII. 3), which is implanted into the tubercle of the radius a little below its neck (fig. LXXVIII. 3). It is an exceedingly thick and powerful muscle, and its manifest action is to bend the fore-arm with great strength. But since its tendon is inserted into the radius, besides bending the fore-arm, it assists other muscles that also act upon the radius in the performance of a function to be described immediately (168).

Fig. LXXVIII.
View of the flexor muscles of the fore-arm. 1. The anterior
surface of the scapula; 2. the muscle called biceps;
3. tendon of the biceps passing to the tubercle of the radius;
4. the muscle called brachialis.

165. The second flexor of the fore-arm, termed the brachialis, is placed immediately under the biceps, and is concealed by it for a considerable part of its course (fig. LXXVIII. 4). Arising from the humerus, on each side of the insertion of the deltoid, it continues its attachment to the bone all the way down the fore part of the humerus, to within inch of the joint; it then passes over the joint, adhering firmly to the anterior ligament (fig. LXXVIII. 4), and is inserted by a strong tendon into the ulna (fig. LXXVIII. 4). It is a thick and fleshy muscle, powerfully assisting the action of the biceps.

166. The two extensor muscles are named the triceps and the anconeous (fig. LXXIX.). The triceps, seated on the back part of the arm, derives its name from having three distinct points of origin, or three separate heads (fig. LXXIX. 5); one of which arises from the scapula and two from the humerus (fig. LXXIX. 5). All these heads adhere firmly to the humerus, as the brachialis does on the fore part of the arm, down to within an inch of the joint (fig. LXXIX. 5), where they form a strong tendon, which is implanted into the olecranon of the ulna (fig. LXXIX. 3); the projection of which affords a lever for increasing the action of the muscle. In all animals that leap and bound, this process of the ulna is increased in length in proportion to their power of performing these movements. The triceps forms an exceedingly thick and strong muscle, which envelops the whole of the back part of the arm (fig. LXXIX.); its action is simple and obvious; it powerfully extends the fore-arm. The anconeous, a small muscle of a triangular form, arising from the external condyle of the humerus, and inserted into the ulna a little below the olecranon, assists the action of the triceps.

Fig. LXXIX.
View of the extensor muscles of the fore-arm. 1. The
scapula; 2. the upper part of the humerus; 3. upper end
of the ulna; 4. upper end of the radius; 5. the muscle
called triceps, the extensor of the fore-arm.

167. Such are the motive powers which act upon the fore-arm, and which produce all the motions of which the hinge-joint of the elbow renders it capable. But besides flexion and extension, the fore-arm is capable of the motion of rotation, which is accomplished by means of the radius. It has been shown (157) that the top of the rounded head of the radius is excavated into a shallow cup (figs. LXIX. 6, and LXXIII. 4) which receives a corresponding convexity of the humerus (figs. LXIX. 2, and LXXIII. 2). In consequence of this articulation with the humerus, the radius, like the ulna, can move backwards and forwards in flexion and extension, the proper movements of the hinge-joint; but that portion of the margin of the hinge of the radius which is in apposition with the ulna is convex (fig. LXIX. 6), and is received into a semilunar cavity hollowed out in the ulna (fig. LXIX. 5). In this cavity the rounded head of the radius revolves, the two bones being held together by the ligament already described (160), which surrounds the head of the radius (fig. LXXVI. 3), and which holds it firmly without being adherent to it, and without impeding in any degree the rotatory motion of the radius. Below, the surface of the radius next the ulna is hollowed out into a semilunar cavity (fig. LXXXII. 1), which receives a corresponding convex surface of the ulna (fig. LXXXII. 2), upon which convex surface the radius rolls (fig. LXXXII. 1). Thus, by the mode in which it is articulated with the ulna above, the radius turns upon its own axis. By the mode in which it is articulated with the ulna below, the radius revolves upon the head of the ulna; and, in consequence of both articulations, is capable of performing the motion of rotation. Moreover, the hand being attached to the radius through the medium of the wrist (figs. LXXXII. 1. 4. and LXXXIII. 1. 4) must necessarily follow every movement of the radius; the rotation of which brings the hand into two opposite positions. In the one, the palm of the hand is directed upwards (fig. LXXXII.); in the other, it is turned downwards (fig. LXXXIII.). When the hand is turned upwards, it is said to be in the state of supination (fig. LXXXII.); when downwards, in that of pronation (fig. LXXXIII.). A distinct apparatus of muscles is provided for effecting the rotation of the radius, in order to bring the hand into these opposite states: one set for producing its supination, and another its pronation.

168. The principal supinators arise from the external condyle of the humerus (fig. LXXX.), and are called long and short (fig. LXXX. 4, 5). The long supinator extends as far as the lower end of the radius, into which it is inserted (fig. LXXX. 4): the short supinator surrounds the upper part of the radius, and is attached to it in this situation (fig. LXXX. 5.). Moreover, the triceps, being inserted into the radius (164), often cooperates with the supinators and powerfully assists their action.

169. The principal pronators are also two, called the round and the square (figs. LXXXI. and LXXXVI. 1). The round pronator arises from the internal condyle, and passing downwards, is inserted into the middle of the radius (fig. LXXXI. 4); the square pronator is a small muscle between the radius and ulna, at their lower extremities being attached to each (fig. LXXXVI. 1).

Fig. LXXX.
View of the supinators of the radius and hand. 1. The
humerus; 2. the ulna; 3. the radius; 4. the muscle called
the long supinator passing to be inserted into the lower portion
of the radius; 5. the muscle, called the short supinator,
surrounding the upper part of the radius.

170. The action of these muscles in producing the rotation of the radius, and so rendering the hand supine or prone, is sufficiently manifest from the mere inspection of the diagrams (fig. LXXXI. 4).

Fig. LXXXI.
View of the pronators of the hand. 1. Lower end of the
humerus; 2. the radius; 3. the ulna; 4. the muscle called
the round pronator, one of the powerful pronators of the hand.

171. The hand is composed of the carpus, metacarpus, and fingers.

172. The carpus (fig. LXXXII. 4) consists of eight small wedge-shaped bones, placed in a double row, each row containing an equal number, and the whole disposed like stones in an arch (fig. LXXXII. 4). They do in fact form an arch, the convexity of which is upwards, on the dorsal surface (fig. LXXXIII. 4); and the concavity downwards, on the palmar surface (fig. LXXXII. 4). But they differ from the stones of an arch in this, that each bone is joined to its fellow by a distinct moveable joint, each being covered with a smooth articulating cartilage. At the same time all of them are tied together by ligaments of prodigious strength, which cross each other in every direction (fig. LXXVII. 6), so that the several separate joints are consolidated into one great joint. The consequence of this mechanism is that some degree of motion is capable of taking place between the several bones, which, when multiplied together, gives to the two rows of bones such an extent of motion, that when the wrist is bent the arch of the carpus forms a kind of knuckle. By this construction a facility and ease of motion, and a power of accommodation to motion and force, are obtained, such as belong to no arch contrived by human ingenuity.

Fig. LXXXII
1. Lower extremity of the radius; 2. lower extremity of
the ulna; 3. styloid process of the ulna; 4. bones of the
carpus or wrist; 5. metacarpal bones; 6. first phalanges
of the fingers; 7. second phalanges of the fingers; 8. third
phalanges of the fingers.

173. The metacarpus (fig. LXXXII. 5), the middle portion of the hand, interposed between the wrist and the fingers, is composed of five bones, which are placed parallel to each other (fig. LXXXII. 5). They are convex outwardly, forming the back (fig. LXXXIII. 5), and concave inwardly, forming the hollow of the hand (fig. LXXXII. 5). They are large at each end, to form the joints by which they are connected with the wrist and fingers (figs. LXXXII. and LXXXIII.): they are small in the middle, in order to afford room for the lodgment and arrangement of the muscles, that move the fingers from side to side (fig. LXXXVI. 2). Their ends, which are joined to the carpus, are connected by nearly plane surfaces (figs. LXXXII. and LXXXIII.): their ends, which support the fingers, are formed into rounded heads, which are received into corresponding cup-shaped cavities, excavated in the top of the first bones of the fingers (fig. LXXXII. 5.). The powerful ligaments that unite these bones pass, both on the dorsal and the palmar surface, from the inferior extremity of the second row of the carpal to the bases of the metacarpal bones (fig. LXXVII, 7, 8). The ligaments are arranged in such a manner as to limit the motions of the joints chiefly to those of flexion and extension, allowing, however, a slight degree of motion from side to side.

174. Each of the fingers is composed of three separate pieces of bone, called phalanges; the thumb has only two (fig. LXXXII. 6, 7, 8): the phalanges are convex outwardly (fig. LXXXII. 6, 7, 8) for increasing their strength, and flattened inwardly (fig. LXXXIII. 6, 7, 8) for the convenience of grasping. The last bones of the fingers, which are small, terminate at their under ends, in a somewhat rounded and rough surface (fig. LXXXIII. 8), on which rests the vascular, pulpy, and nervous substance, constituting the special organ of touch, placed at the points of the fingers, and guarded on the upper surface by the nail (fig. LXXXII. 8).

Fig. LXXXIII.
1. Lower extremity of the radius; 2. lower extremity of
the ulna; 3. styloid process of the ulna; 4. bones of the
carpus; 5. metacarpal bones; 6. 7. 8. first, second, and
third phalanges of the fingers.

175. The round inferior extremity of the metacarpus is admitted into the cavity of the superior extremity of the first phalanx of the five fingers (figs. LXXXII. and LXXXIII.), and their joints are connected by lateral and transverse ligaments of great strength (fig. LXXVII. 9). The situation and direction of the ligaments which unite the several phalanges of the fingers (fig. LXXVII. 9) are precisely the same as those of the articulation of the phalanges with the metacarpus (fig. LXXVII. 7, 8); and the articulation of these bones with one another is such as to admit only of the motions of flexion and extension.

176. The muscles which perform these motions are seated for the most part on the fore-arm. Independently of the supinators and pronators which have been already described (167 et seq.), there are distinct sets of muscles for bending and extending the wrist and the fingers. The flexors arise from the internal, and the extensors from the external, condyle of the humerus (fig. LXIX. 3, 4). The internal condyle is larger and longer than the external (fig. LXIX. 3, 4); for the flexors require a larger point of origin and a longer fulcrum than the extensor muscles; because to the actions of flexion, such as grasping, bending, pulling, more power is necessary than to the action of extension, which consists merely in the unfolding or the opening of the hand previously to the renewal of the grasp.

177. For the same reason, two muscles are provided for flexing, while only one is provided for extending the fingers. The flexors, bulky, thick, and strong, are placed on the fore part of the fore-arm (fig. LXXXIV.). The first, named the superficial flexor (fig. LXXXIV. 1), about the middle of the arm, divides into four fleshy portions, each of which ends in a slender tendon (fig. LXXXIV. 1). As these tendons approach the fingers they expand (fig. LXXXIV. 1), and when in apposition with the first phalanx, split and form distinct sheaths for the reception of the tendons of the second flexor (fig. LXXXIV. 3). After completing the sheath, the tendons proceed forward along the second phalanx, into the fore part of which they are implanted, and the chief office of this powerful muscle is to bend the second joint of the fingers upon the first, and the first upon the metacarpal bone. Its action is assisted by a second muscle, called the deep or profound flexor (fig. LXXXIV. 2), because it lies beneath the former; or the perforans, because it pierces it. Bulky and fleshy, this second flexor, like the first, about the middle of the arm, divides into four tendons, which, entering the sheaths prepared for them in the former muscle (where the tendons are small and rounded for their easy transmission and play), pass to the root of the third phalanx of the fingers into which they are implanted (fig. LXXXIV. 3).

Fig. LXXXIV.
View of the flexor muscles of the fingers. 1. The superficial
flexor, divided and turned aside, to show, 2. the deep
flexor; 3. sheaths for the tendons of the deep flexor,
formed by the splitting of the tendons of the superficial
flexor; 4. the anterior annular ligament, divided and turned
aside.

118. The muscle that extends the fingers, called the common extensor, is placed on the back part of the fore-arm (fig. LXXXV.), about the middle of which it divides into four portions which terminate in so many tendons (fig. LXXXV. 2). When they reach the back of the metacarpal bones, these tendons become broad and flat, and send tendinous expansions to each other, forming a strong tendinous sheath which surrounds the back of the fingers (fig. LXXXV. 2). These tendinous expansions are inserted into the posterior part of the bones of the four fingers (fig. LXXXV. 2); and their office is powerfully to extend all the joints of all the fingers (fig. LXXXV. 2).

179. On both the palmar and dorsal regions of the wrist are placed ligaments for tying down these tendons, and preventing them from starting from their situation during the action of the muscles (figs. LXXXIV. and LXXXV.). On the palmar region an exceedingly strong ligament passes anteriorly to the concave arch of the carpus (fig. LXXXIV. 4) for the purpose of tying down the tendons of the flexor muscles. On the dorsal surface (fig. LXXXV.), a similar ligament, passing in an oblique direction from the styloid process of the radius to the styloid process of the ulna (fig. LXXXV. 3), performs the same office in tying down the tendons of the extensor muscle. Both these ligaments are called annular.

Fig. LXXXV.
View of the extensor muscles of the fingers. 1. The
common extensor, sending (2 2 2 2) tendons to each
finger; 3. the posterior annular ligament.

180. In the palm of the hand are placed additional muscles which assist the flexors of the fingers (fig. LXXXVI. 2), being chiefly useful in enabling the fingers to perform with strength and precision short and quick motions. There are especially four small and rounded muscles (fig. LXXXVI. 2), resembling the earth worm in form and size, and hence called lumbricales; but as their chief use is to assist the fingers in executing short and rapid motions, they have also received the better name of the musculi fidicinales.

Fig. LXXXVI.
1. The muscle called the square pronator; 2. muscles
seated in the palm of the hand, by which, chiefly, the
fingers execute short and rapid motions.

181. The thumb, in consequence of the comparative looseness of its ligaments, is capable of a much greater extent of motion than the fingers, and can be applied to any part of each of the fingers, to different parts of the hand, and in direct opposition to the power exerted by the whole of the fingers and hand, in the act of grasping. The muscles which enable it to perform these varied motions, and which act powerfully in almost every thing we do with the hand, form a mass of flesh at the ball of the thumb (fig. LXXXVII. 1), almost entirely surrounding it. The little finger is also provided with a distinct apparatus of muscles (fig. LXXXVII. 2), which surrounds its root, just as those of the thumb surround its ball, in order to keep it firm in opposition to the power of the thumb in the act of grasping, and in various other motions.

Fig. LXXXVII.
1. The mass of muscles forming the ball of the thumb;
2. the mass of muscles forming the ball of the little finger;
3. tendons of one of the flexor muscles of the fingers;
4. sheaths formed by the tendons of the superficial flexor
for the reception of the tendons of the deep flexor.

182. The upper extremity is covered by a tendinous expansion or fascia which envelopes the whole arm, encloses its muscles as in a sheath, and affords them, in their strong actions, "that kind of support which workmen feel in binding their arms with thongs." This fascia likewise descends between many of the muscles, forming strong partitions between them, and affording points of origin to many of their fibres, scarcely less fixed than bone itself.

183. From the whole, it appears, that the first joint of the upper extremities, that of the shoulder, is a ball and socket joint, a joint admitting of motion in every direction; that the second joint, that of the elbow, is partly a hinge-joint, admitting of flexion and extension, and partly a rotation joint, admitting of a turning or rotatory motion; and that the joints of the wrist and of the fingers are likewise hinge-joints, admitting at the same time of some degree of lateral motion. When these various motions are combined, the result is that the hand can apply itself to bodies in almost every direction, in any part of the area described by the arm, when all the joints are moved to their utmost extent. There is thus formed an instrument of considerable strength, capable of a surprising variety and complexity of movements, capable of seizing, holding, pulling, pushing and striking with great power, yet at the same time capable of apprehending the minutest objects, and of guiding them with the utmost gentleness, precision, and accuracy, so that there are few conceptions of the designing mind which cannot be executed by the skilful hand.

184. The lower extremities consist of the thigh, leg, and foot.

185. The osseous part of the thigh consists of a single bone, called the femur (fig. XXXIV. 4), the longest, thickest, and strongest bone in the body. It sustains the entire weight of the trunk, and occasionally much heavier loads superimposed upon it. It is constructed in such a manner as to combine strength with lightness. This is effected by rendering the bone what is technically called cylindrical; that is, a bone in which the osseous fibres are arranged around a hollow cylinder. There are two varieties of osseous matter,—the compact, in which the fibres are dense and solid (fig. LXXXVIII. 1), and the spongy, in which the fibres are comparatively tender and delicate (fig. LXXXVIII. 2). Both varieties are, indeed, combined, more or less, in every bone, the compact substance being always external, and the spongy internal; but in the cylindrical bones the arrangement is peculiar. Every long or cylindrical bone consists of a body or shaft (fig. LXXXVIII. 4.), and of two extremities (fig. LXXXVIII. 5). The body is composed principally of compact substance, which on the external surface is so dense and solid, that scarcely any distinct arrangement is visible; but towards the interior this density diminishes; the fibres become distinct (fig. LXXXVIII. 5), and form an expanded tissue of a cellular appearance (fig. LXXXVIII. 5), the cells being called cancelli, and the structure cancellated. In the centre of the bone even the cancelli disappear; the osseous fibres terminate; and a hollow space is left filled up, in the natural state, by an infinite number of minute membranous bags which contain the marrow (fig. LXXXVIII. 3). In the body of the bone, to which strength is requisite, that part being the most exposed to external violence, the compact matter is arranged around a central cavity. By this means strength is secured without any addition of weight; for the resisting power of a cylindrical body increases in proportion to its diameter; consequently the same number of osseous fibres placed around the circumference of a circle produce a stronger bone than could have been constructed had the fibres been consolidated in the centre, and had the diameter been proportionally diminished. The hollow space thus gained in its centre, renders the bone lighter by the subtraction of the weight of as many fibres as would have gone to fill up that space; while its strength is not only not diminished by this arrangement, but positively increased. On the other hand, at the extremities of the bone, space, not strength, is required; required for the attachment and arrangement of the tendons of the muscles that act upon it, and for the formation of joints (fig. LXXXVIII. 5). Accordingly, at its extremities the bone swells out into bulky surfaces; but these surfaces are composed, not of dense and solid substance, but of spongy tissue, covered by an exceedingly thin crust of compact matter, and so, as by the former expedient strength is secured without increase of weight, by this, space is obtained without increase of weight.

Fig. LXXXVIII.
A section of the femur, showing, 1. the compact bony
substance; 2. the spongy or cancellated structure; 3. the
internal cavity containing the marrow; 4. body; 5, extremities
of the bone.

186. The thigh-bone, placed at the under and outer part of the pelvis, has an oblique direction, the under being considerably nearer its fellow than the upper end (fig. XXXIV. 4), in order to afford space for the passages at the bottom of the pelvis, and also to favour the action of walking. The body of the bone, which is of a rounded form (fig. XXXIV. 4), is smooth on its anterior surface (fig. XXXIV. 4), where it is always slightly convex, the convexity being forwards (fig. XXXIV. 4), while its posterior surface is irregular and rough, and forms a sharp prominent line, termed the linea aspera (fig. XXXV. 4), giving attachment to numerous muscles.

187. The superior extremity of the femur terminates in a large ball or head, which forms nearly two-thirds of a sphere (fig. LXXXIX. 4.). It is smooth, covered with cartilage, and received into the socket of the ilium called the acetabulum, which, deep as it is, is still further deepened by the cartilage which borders the brim (fig. LXXXIX. 3). The brim is particularly high in the upper and outer part, because it is in this direction that the reaction of the ground against the descending weight of the trunk tends to dislodge the ball from its socket.

188. Passing obliquely downwards and outwards from the ball, is that part of the femur which is called the neck (fig. LXXXIX. 5). It spreads out archlike between the head and the body of the bone, and is more than an inch in length (fig. LXXXIX. 5). It is thus long in order that the head of the bone may be set deep in its socket, and that its motions may be wide, free, and unembarrassed.

Fig. LXXXIX.
1. Lower portion of the ilium; 2. tuberosity of the
ischium: 3. socket for the head of the femur, or thigh-bone;
4. head of the femur; 5. neck of the femur; 6. the
great process of the femur called the trochanter major;
7. the body of the femur.

189. From the external surface of the femur, nearly in a line with its axis, proceeds the largest and strongest bony process of the body which gives insertion to its most powerful muscles, namely, those that extend the thigh and that turn it upon its axis (fig. LXXXIX. 6). Because, from its oblique direction, it rotates the thigh, this process is called the trochanter, and, from its size, the trochanter major. At the under and inner part of the neck on the posterior surface of the bone, is a similar process, but much smaller, called the trochanter minor (fig. XXXV. 4), into which are inserted the muscles that bend the thigh.

190. The inferior extremity of the femur, much broader and thicker than the superior (fig. XC. 1), is terminated by two eminences, with smooth surfaces, termed condyles (fig. XC. 2), which, articulated with the tibia, and the patella, form the joint of the knee (figs. XC. 2, 4, 5, and XCI. 1, 2, 3).

Fig. XC.
1. Lower end of the femur; 2. condyles of the femur;
3. upper end of the tibia; 4. articular surfaces on the head
of the tibia on which the thigh-bone plays; 5. the patella,
or knee-pan; 6. upper end of the fibula, not entering into
the knee-joint.

Fig. XCI.
Posterior view of the bones forming the knee-joint.
1. Lower end of the femur; 2. upper end of the tibia;
3. articular surfaces on the head of the tibia, on which
the thigh-bone plays; 4. upper end of the fibula, not entering
into the knee joint.

191. The bones of the leg, two in number, consist of the tibia (fig. XC. 3) and fibula (fig. XC. 6). The tibia, next to the femur, the longest bone in the body, is situated at the inner side of the leg (fig. XC. 3). Its superior extremity is bulky and thick (fig. XC. 3). The top of it forms two smooth and slightly concave surfaces, adapted to the convex surfaces of the condyles of the femur (fig. XC. 4, 2). On its outer side there is a smooth surface, to which the head of the fibula is attached (fig. XC. 6). Its lower extremity, which is small, forms a concavity adapted to the convexity of the bone of the tarsus, called the astragalus, with which it is articulated (fig. XCII. 4.) Its inner part is produced so as to form the inner ankle (figs. XCII. 2, and XCIII. 3): its outer side is excavated into a semilunar cavity, for receiving the under end of the fibula, which forms the outer ankle (figs. XCII. 3, and XCIII. 4).

192. The fibula, in proportion to its length the most slender bone of the body, is situated at the outer side of the tibia (fig. XC. 6). Its upper end formed into a head, with a flat surface on its inner side (figs. XC. 6, and XCI. 4), is firmly united to the tibia (fig. XC. 4). Its lower end forms the outer ankle, which is lower and farther back than the inner (fig. XCII. 3, 2).

Fig. XCII.
Anterior view of the bones forming the ankle-joint.
1. Lower end of the tibia; 2. production of the tibia, forming
the inner ankle; 3. lower end of the fibula, forming the
outer ankle; 4. upper part of the astragalus: these three
bones form the ankle-joint; 5 5 5, other bones of the tarsus;
6 6 6 6 6 metatarsal bones.

Fig. XCIII.
Posterior view of the bones forming the ankle-joint.
1. Lower end of the tibia; 2. lower end of the fibula; 3. internal
malleolus or ankle; 4. external malleolus or ankle;
5. one of the tarsal bones, called the astragalus, with which
the tibia and fibula are articulated; 6. the os calcis or heel.

193. The patella, or knee-pan (fig. XC. 5), is a light but strong bone, of the figure of the heart as painted on playing-cards, placed at the fore part of the joint of the knee, and attached by a strong ligament to the tibia, the motions of which it follows (fig. XC. 5). It is lodged, when the knee is extended, in a cavity formed for it in the femur (fig. XC.); when bent, in a cavity formed for it at the fore part of the knee (fig. XC. 5).

194. The foot consists of the tarsus, metatarsus, and toes.

195. The tarsus, or instep, is composed of seven strong, irregular-shaped bones, disposed like those of the carpus, in a double row (fig. XCII. 4, 5). The arrangement of the tarsal bones is such as to form an arch, the convexity of which above, constitutes the upper surface of the instep (fig. XCII. 4, 5): in the concavity below are lodged the muscles, vessels, and nerves that belong to the sole.

196. The metatarsus consists of five bones, which are placed parallel to each other (fig. XCII. 6), and which extend between the tarsus and the proper bones of the toes (fig. XCII. 6). Their extremities, especially next the tarsus, are large, in order that they may form secure articulations with the tarsal bones (fig. XCII. 6). Their bodies are arched upwards (fig. XCII. 6), slightly concave below, and terminate forwards in small, neat, round heads, which receive the first bones of the toes, and with which they form joints, admitting of a much greater degree of rotation than is ever actually exercised, in consequence of the practice of wearing shoes. The natural, free, wide-spreading form of the toes, and the consequent security with which they grasp the ground, is greatly impaired by this custom. Taken together, the bones of the metatarsus form a second arch corresponding to that of the tarsus (fig. XCVIII. 2).

197. Each toe consists of three distinct bones, called, like those of the fingers, phalanges (fig. XCVIII.), but the great toe, like the thumb, has only two (fig. XCVIII.). That extremity of the first phalanges which is next the metatarsal bones is hollowed into a socket for the head of the metatarsal bones.

198. Besides the bones already described, there are other small bones, of the size and figure of flattened peas, found in certain parts of the extremities, never in the trunk, called sesamoid, from their resemblance to the seed of the sesamum. They belong rather to the tendons of the muscles than to the bones of the skeleton. They are embedded within the substance of tendons, are found especially at the roots of the thumb and of the great toe, and are always placed in the direction of flexion. Their office, like that of the patella, which is, in truth, a bone of this class, is to increase the power of the flexor muscles by altering the line of their direction, that is, by removing them farther from the axis of the bone on which they are intended to act.

199. The ligaments which connect the bones of the lower extremities are the firmest and strongest in the body. Of these, the fibrous capsule of the hip-joint (fig. XCIV. 1), which secures the head of the femur in the cavity of the acetabulum (fig. XCIV.), is the thickest and strongest. It completely surrounds the joint (fig. XCIV. 1). It arises from the whole circumference of the acetabulum, and, proceeding in a direction outwards and backwards, is attached below to the neck of the femur (fig. XCIV. 1). It is thicker, stronger, and much more closely attached to the bones than the fibrous capsule of the shoulder-joint (144), because the hip-joint is formed, not like the shoulder-joint, for extent of motion, but for strength. Its internal surface is lined by synovial membrane, and its external surface is covered and strengthened by the insertion of muscles that move the thigh-bone. The joint is strengthened by another ligament, which passes from the inner and fore part of the cavity of the acetabulum (fig. XCV.) to be inserted into the head of the femur (fig. XCIV.), called the round ligament, the office of which obviously is to hold the head of the femur firmly in its socket.

Fig. XCIV.
1. The fibrous capsule of the hip-joint, laid open and
turned aside to show, 2. the round ligament in its natural
position.
Fig. XCV.
A view of the head of the femur drawn out of its socket,
and suspended by the round ligament, to show more clearly
the action of the ligament in retaining the head of the
femur in its socket.

200. Numerous and complicated ligaments connect the bones that form the knee-joint (fig. XCVI.), and the strength of these powerful bands is greatly increased by the tendons that move the leg (fig. XCVI. 5), which pass over, and more or less surround, the joint.

Fig. XCVI.
General view of the ligaments of the knee-joint. 1. Lower
end of the femur; 2. upper end of the tibia; 3. upper end
of the fibula; 4. the patella; 5. united tendons of the
extensor muscles; 6. ligaments of the patella; 7. the capsular
investment of the knee; 8. the internal lateral ligament;
9. the external lateral ligaments; 10. the posterior
ligament; 11. the ligament connecting the tibia and fibula;
12. a portion of the interosseous ligament.

201. Strong ligaments maintain in their proper position the bones that form the ankle-joint (fig. XCVII.), connect the bones of the tarsus and metatarsus with one another (fig. XCVIII. 1), and articulate the several phalanges of the toes (fig. XCVIII. 2).

Fig. XCVII.
General view of the posterior ligaments of the ankle-joint.
1. Lower end of the tibia; 2. lower end of the
fibula; 3. astragalus; 4. os calcis; 5. ligament between
the tibia and fibula; 6. ligament passing from the fibula
to the astragalus; 7. ligament passing from the fibula to
the os calcis; 8. ligament passing from the tibia to the
astragalus.

Fig. XCVIII.
General view of the ligaments of the sole of the foot.
1. Ligaments connecting the bones of the tarsus; 2. ligaments
connecting the bones of the toes.

202. The joint of the hip, like that of the shoulder, is capable of flexion, extension, and rotation; but its rotatory motions are to a much less extent, on account of the greater depth of the acetabulum and the stronger and shorter fibrous capsule. When the femur is flexed, the thigh is bent upon the pelvis, and its inferior extremity is carried forwards. When it is extended, the thigh is carried backwards. The two thighs may be separated from each other laterally (abduction), or brought near to each other (adduction), or the one may be made to cross the other, and they may be rotated outwards or inwards.

203. The apparatus of muscles that produces these varied motions is seated partly on the trunk and partly on the pelvis. Thus, the powerful muscle that flexes the thigh, or that carries it forwards, termed the psoas (fig. XCIX. 1), arises from the last vertebra of the back, and successively from each vertebra of the loins (fig. XCIX. 1), and is inserted into the lesser trochanter of the femur (fig. XCIX. 3). Its action is assisted first by a large and strong muscle named the iliacus (fig. XCIX. 2), which occupies the whole concavity of the ilium (fig. XCIX. 2), and which, like the psoas, is inserted into the lesser trochanter of the femur (fig. XCIX. 3).

Fig. XCIX.
View of the muscles that bend the thigh. 1. The
muscle called psoas; 2. the muscle called iliacus; 3. tendons
of these muscles, going to be inserted into the trochanter
minor of the femur.

204. The muscles that extend the thigh, or that carry it backwards, named the glutÆi, the most powerful muscles of the body, are placed in successive layers, one upon the other, on the back part of the ilium (fig. C. 1, 2, 3), and are inserted into the linea aspera of the femur. They constitute the mass of flesh which forms the hip, and their powerful action in drawing the thigh backwards is assisted by several other muscles (fig. C. 4, 5, 6). Their action is never perfectly simple and direct; for those which move the thigh forwards sometimes carry it inwards, and sometimes outwards; and in like manner, those which move it backwards, at one time carry it inwards and at another outwards, according to the direction of the fibres of the muscle and the position of the limb when those fibres act; while some of them, and more especially those which carry it backwards, at the same time rotate it, or roll it upon its axis.

Fig. C.
View of the muscles that extend the thigh. 1. The
muscle called glutÆus maximus, removed from its origin,
2, 2, to show the muscles which lie beneath it; 2. cut edge
showing the origin of the same muscle; 3. the muscle
called glutÆus medius; 4, 5, 6. smaller muscles, assisting
the action of the glutÆi.

205. The knee is a hinge-joint, admitting only of flexion and extension, and is therefore provided only with two sets of muscles, one for bending and the other for extending the leg. The flexors of the leg arise from the under and back part of the pelvis, are seated on the back part of the thigh, and are inserted into the upper part either of the tibia or of the fibula (fig. CI). They consist for the most part of three muscles, named the semi-tendinosus, the semi-membranosus (fig. CI. 3), and the biceps of the leg (fig. CI. 1). The tendons of the two former muscles, in passing to be inserted into the leg, form the inner, and that of the latter the outer, hamstrings (fig. CI. 4, 5).

206. Four large muscles, blended together in such a manner as to form one muscle of prodigious size, termed the quadriceps cruris (fig. CI. 7), occupying nearly all the forepart and the sides, and a considerable portion of the back part of the thigh, constitute the great flexor of the thigh. This enormous mass of muscle arises partly from the ischium, and partly from the upper part of the femur (fig. CI. 7), and is all inserted into the patella (fig. CI. 8), which constitutes a pulley for the purpose of assisting the action of these powerful muscles.

207. The muscles which bend the toes and extend the foot, termed the gastrocnemii (fig. CII. 1, 2), are placed on the back part of the leg, and form the mass of muscle which constitutes the calf of the leg (fig. CII. 1, 2). They arise partly from the lower extremity of the femur (fig. CII.) and partly from the upper and back part of the fibula and tibia; and they form the largest and strongest tendon in the body, termed the tendo achillis (fig. CII. 3), which is implanted into the heel (fig. CII. 4).

Fig. CII.
View of the muscles which bend the toes, and which, by
lifting the heel, extend the foot. 1. The muscle called
gastrocnemius externus, which, uniting with 2. the gastrocnemius
internus, forms 3. the tendo achillis, which is inserted
into 4. the heel.

Fig. CIII.
View of the muscles which extend the toes and bend the
foot. 1. The common extensor; 2. the tendons of the same
muscle inserted into the toes; 3. the anterior annular
ligament of the foot.

Fig. CIV.
View of the muscles in the sole of the foot. 1 The
muscle which draws the great toe from the other toes; 2. the
muscle which draws the little toe from the other toes;
3. the muscle called the short flexor of the toes, which
assists in bending the four smaller toes.

208. The muscles which extend the toes and bend the foot are seated on the fore part of the leg (fig. CIII.); split into tendons like the analogous muscles of the fingers (fig. CIII. 2); and are bound down by a ligament (fig. CIII. 3), exactly the same in name, disposition, and office, as that which belongs to the hand (fig. CIII. 3). Numerous minute muscles are placed in the sole of the foot (fig. CIV.), which act on the toes as the small muscles in the palm of the hand act on the fingers (fig. LXXXVI.).

209. Such are the moving powers which put in action the complicated mechanism provided for the function of locomotion. And these powers are adequate to their office; but they are what may be termed expensive powers; agents requiring a high degree, of organization and the utmost resources of the economy to support and maintain them. Hence in the construction of the framework of the machine which they have to move, whatever mechanical contrivance may economize their labour, is adopted. The construction, form, and disposition of the several parts of that framework have all reference to two objects: first, the combination of strength with lightness; and secondly, security to tender organs, with the power of executing rapid, energetic, and, sometimes, violent motions. The combination is effected and the object attained in a mode complicated in the detail, simple in the design, and perfect in the result. The weight of the body transmitted from the arch of the pelvis to a second arch, formed by the neck of the thigh-bone, and from this, in a perpendicular direction, to a third arch formed by the foot, is ultimately received by the heel behind, and by the metatarsal bones and the first phalanges of the toes before, and more especially by the metatarsal joints belonging to the great and little toe, which have a special apparatus of muscles, for the purpose of preserving steadily their relative situation to the heel. The weight of the body is thus sustained on a series of arches, from which it is, in succession, transmitted to the ground, where it ultimately rests upon a tripod: forms known and selected as the best adapted to afford support, and to give security of position. Columns of compact bone superimposed one upon another, and united at different points by bands of prodigious strength, form the pillars of support. But these bony columns never touch each other; are never in actual contact; are all separated by layers of elastic matter which, while they assist in binding the columns together, enable them to move one upon another, as upon so many pliant springs. The layers of cartilage interposed between the several vertebrÆ; the layer of cartilage interposed between the vertebral column and the pelvis; the layer of cartilage that lines the acetabulum and that covers the head of the femur; the layer of cartilage that covers the lower extremity of the femur and the upper extremity of the tibia and fibula and the tarsus; the successive layers of cartilage interposed between the several bones of the tarsus; and finally, the layer of cartilage that covers both the tarsal and the digital extremities of the metatarsal bones; are so many special provisions to prevent the weight of the body from being transmitted to the ground with a shock; and, at the same time, so many barriers established between the ground and the spinal cord, the brain and the soft and tender organs contained in the thoracic and abdominal cavities, to prevent these organs from being injured by the reaction of the ground upon the body. The excellence of this mechanism is seen in its results; in contemplating "from what heights we can leap—to what heights we can spring—to what distances we can bound—how swiftly we can run—how firmly we can stand—how nimbly we can dance—and yet how perfectly we can balance ourselves upon the smallest surfaces of support!"

210. It is necessary, in order to complete this general view of the structure of the human body, and of the combination and arrangement of its various parts, to denote the several regions into which, for the purpose of describing with accuracy the situation and relation of its more important organs, the body is divided. It is not needful to the present purpose to describe the regions of the head, because its internal cavity contains only one organ, the brain, and its external divisions do not differ materially from those which are common and familiar; but the chest, the abdomen, and the upper and lower extremities are mapped out into regions, of which it is very important to have an exact knowledge, which may be acquired by the study of the annexed diagrams.

Fig. CV.
Anterior view of the regions of the body. 1. Region of the neck;
2. region of the chest or thorax. Abdominal regions: 3. epigastric;
4. umbilical; 5. hypogastric region. Regions of the upper extremities.
6. shoulder; 7. arm; 8. elbow; 9. fore-arm; 10. wrist; 11. ball of
thumb; 12. the axilla or armpit. Regions of the lower extremities:
13. thigh; 14. knee; 15. leg; 16. ankle; 17. instep and foot.

Fig. CVI.
Posterior view of the regions of the body: 18. region to
the scapula; 19. of the back; 20. of the loins; 21. of the
hips; 22. of the ham; 23. of the calf of the leg; 24. of the
heel and foot.

Fig. CVII.
Lateral view of the regions of the body: 25. arch of
the foot.

Fig. CVIII.
Anterior view of the situation of the more important internal
organs: 1. lungs, right and left; 2. heart; 3. line
representing the edge of the diaphragm; 4. liver; 5. stomach;
6. small intestines; 7. colon; 8. urinary bladder.

Fig. CIX.
Posterior view of the situation of the more important
internal organs: 9. kidnies, right and left; 10. the course
of the spinal cord.

Fig. CX.
Lateral view of the situation of the more important
internal organs.

                                                                                                                                                                                                                                                                                                           

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