PLANT NOTES

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THE FERNS. The Sliding Sands Trail drops from White Hill on bare slopes of red and gray Cinders. As it levels below Puu o Pele, a lush, green carpet spreads along the south wall of the crater. It is a surprise to discover that ferns compose the verdure, for several kinds find the shelter of the cliffs agreeable. Haleakala’s KA’UPU, Polystichum haleakalense Brack., a rather coarse, low plant with scaly stems, grows among the shrubs. The pellucid polypody, ’AE, Polypodium pellucidum Kaulf., and the maidenhair spleenwort, ’IWA’IWA, Asplenium adiantum-nigrum L., grow here, but they are common elsewhere as well. Iwaiwa is a small plant with shiny, slender stems and stiff, triangular fronds that thrives in the brush. On Leleiwi Pali it is a tiny thing, clinging to the rocks. This fern is known in mountainous regions in many countries. Another maidenhair spleenwort, ’OWALI’I, Asplenium trichomanes L., makes its home on barren lava above 5,000 feet on the inner slopes of the crater. Its small, opposite, rounded or ovate frond segments, pinnae, grow on wiry stems that form dense clumps. Look for it along Halemauu Trail on Leleiwi Pali. It is found in the temperate zones and on high mountains in many parts of the world.

Bracken, KILAU, Pteridium aquilinum var. decompositum (Gaud.) Tryon, a stiff cosmopolitan, is among the best-known ferns. It grows in grassy spots inside and outside the crater. Foraging pigs, seeking its tasty rootstocks, often uproot it in the forests. The young leaves of AMA’UMA’U, Sadleria cyatheoides Kaulf., add a touch of red along Halemauu Trail on Leleiwi Pali. At Paliku, several moisture-loving ferns grow on logs, rocks, and moss-covered trees. Large, dense ae are quite unlike the frail specimens on Leleiwi Pali. The following can be seen in damp spots and on tree trunks: Maui’s paddle, ’EKAHA, Elaphoglossum reticulatum (Kaulf.) Gaud., with paddle-shaped blades 6-12 inches long; ’EKAHA-’AKOLEA, Pleopeltis thunbergiana Kaulf., a small fern with tough, pale, elliptico-oblong (paddle-shaped) fronds; and MOA, Psilotum complanatum Sw., belonging to a small group of tropical plants, PSILOTALES, which reproduce by spores, but are distinct from true ferns, clubmosses, and the better-known orders.

Swordfern, NI’ANI’AU or ’OKUPUKUPU, Nephrolepis exaltata (L.) Schott., is widely distributed at Haleakala as it is elsewhere in Hawaii. Cliffbrake, KALAMOHO LAULI’I, Pellaea ternifolia (Cav.) Link, grows among rocks in dry, sunny locations at higher elevations above the park entrance and inside the crater. It is common. The bluish-green pinnae of this short, slender fern grow on opposite sides of the dark, wiry stems. They are cleft into three linear segments.

THE NATIVE GRASSES. Several native grasses grow above the park entrance: Trisetum glomeratum (Kunth) Trin., Deschampsia australis forma haleakalensis Skottsb., and Agrostis sandwicensis Hillebr. The Trisetum, also common inside the crater, is called mountain pili, PILI being the lowland grass known in many tropical regions and used for thatching houses in Hawaii. The Hawaiian name for T. glomeratum is PILIUKA, upland pili. In some places it is also called HE’U PUEO, the hoot of an owl.

All three grasses are tufted, i.e., bunch grasses. The Deschampsia has tough, wiry blades (leaves) with shiny, open panicles of flowers and seeds. The Agrostis has stiffly upright blades and culms (jointed stems) with spike-like panicles. The pili has flat blades usually covered with soft hairs. The panicles are contracted or spike-like.

THE SEDGES. Fig. 1. Sedges are generally grasslike wind-pollinated herbs that grow in tufts or bunches. They are often the dominant plants in cold marshes, especially in the Arctic. They have little economic value and grazing animals find them unpalatable. Some have tough, pliable stems that are woven into mats and baskets; some have fruiting spikes that are attractive in dry bouquets. Most have 3-angled stems around which the blades are ranked. The inconspicuous, green flowers are crowded in tight, flattened spikes, often grouped on top of a slender, grasslike stem. Hawaii has a dozen or more native genera with many of its species widespread in the world.

The species common on the upper slopes of Haleakala and on the crater floor, Gahnia gaudichaudii Steud., bears shiny, ebony fruits. These can dangle, suspended by the wilted, threadlike stamens for more than a year from the fruiting stalk. Another sedge, Carex macloviana subfusca (W. Boott) Kukenth., grows in clumps along the south wall. The fruiting stalks bear cylindrical or ovoid clusters of 4-9 spikelets, each ½-1 inch long. This sedge grows from Lapland and Greenland to northern South America, but in Hawaii it has been found only on Haleakala and Kohala Mountains above 4,000 feet. The Hawaiian variety was first described from a specimen collected at Lake Tahoe, California. The interesting Oreobolus furcatus H. Mann, mentioned as occurring on West Maui, also grows in Koolau Gap.

RUSH, Luzula hawaiiensis (O. Ktze.) Buch. A visitor brought me a small tufted plant with grass-like leaves covered with soft, silky hairs. He was all excited, believing he had found a young silversword on the cinder flats near the summit of Mt. Hualalai. The plant was the endemic rush that lives in wet places above 3,000 feet in the mountains of Maui, Hawaii, and Kauai. In Haleakala Crater, I found it on Leleiwi Pali and near Paliku, where others might mistake it for an immature silversword or greensword. The Hawaiian general name for grasses, sedges, and rushes is MAU’U.

PAINIU, Astelia degeneri Skottsb. This plant is reported by Degener to be growing within the crater.[19]

MAU’U-LA’ILI, Sisyrinchium acre H. Mann. A native member of the Iris Family with grass-like leaves that grows on old lava flows between altitudes of 3,500 and 7,500 feet on Maui and Hawaii. It is an attractive plant that rarely exceeds 12 inches high. In July and August it bears yellow flowers ¾ inch across that last only a few hours. In the crater it may be found in Koolau and Kaupo Gaps. The leaves, bound tightly around the wrists and ankles, stain the skin a blue color that lasts several days. This stain was regarded as proof that a person had been to the crater. A number of other species of the genus are mainland wildflowers commonly called blue-eyed grass.

ORCHIDS. Fig. 2. The Orchid Family of over 15,000 species is second only to the Composite Family in size, yet only three species of three genera are native in Hawaii, a land connected in thought with an exuberance of gay orchids. Moreover, the three endemic species are characterized by small size, relative rarity, and inconspicuous flowers. Twayblade, Liparis hawaiiensis H. Mann, grows in open woods on the ground and on moss-covered trees on the flanks of the volcano and at Paliku. The rarest of the three natives, Habenaria holochila Hillebr., has been found growing 1-2½ feet high in deep moss in fog-swept Koolau Gap. It bears its dull-greenish, inconspicuous orchids on a tall, many-flowered spike.

’ALA ’ALAWAINUI, Peperomia sp. A small herb with succulent leaves found at Paliku. It is a member of the Pepper Family, Piperaceae, to which AWA, Piper methysticum, belongs. There are many Peperomias native to Hawaiian forests. Some have leaves with gay red undersides.

SANDALWOOD, ’ILIAHI, Santalum haleakalae Hillebr. Fig. 3. A small but striking tree found above the park entrance, in Koolau Gap, and along the Kaupo Trail below Paliku. It has leathery, dark green leaves so that it stands out in the vegetation and is readily distinguished from afar. In mid-summer, corymbs of four-pointed, deep-red or vermillion flowers appear on the ends of branches. The dry heartwood has the fragrance which is associated with the name sandalwood. Degener tells the detailed story of sandalwood trade that flourished in the islands for fifty years beginning in 1790.[20]

SHEEP SORREL, Rumex acetosella L., an abundant, well-known, introduced weed found both inside the crater and out. A native species called PAWALE, Rumex gigantius Ait., is more interesting. It grows as a stocky undershrub on barren lava flows and in rock crevices inside the crater. In Koolau Gap, it becomes a sprawling vine. It is said that a mixture of an extraction from the boiled bark and AWA was used by Hawaiians for skin diseases. This was tried in vain as a cure for leprosy when the disease first appeared in the islands in 1840.

HAWAIIAN BUTTERCUP, MAKOU, Ranunculus hawaiiensis A. Gray. Like the native violets and geraniums, members of this genus, well-known on the mainland, either become bushy or spread as a woody vine in Hawaii. Two native species occur on Haleakala, but the yellow-flowered, sprawling R. mauiensis A. Gray is reported only outside the park. The larger flowered, erect Hawaiian buttercup, however, was reported by J. F. Rock to be abundant formerly in Puu Nianiau Crater. It is a coarse, hollow-stemmed, hairy herb with compound leaves divided into three sharply-toothed, irregular leaflets. Within Haleakala Crater it grows on moist, grassy slopes in Koolau Gap.

HO’AWA, Pittosporum confertiflorum A. Gray. A small tree, sometimes becoming 25 feet tall. The large leaves, shiny green on top, brown hairy underneath, are crowded like whorls on the ends of the branches. In the centers of these, dense clusters of fleshy, cream-colored flowers appear in late summer. Wrinkled fruits, resembling English walnuts, hang on the trees throughout the year. Very few trees grow in the crater, but they are more numerous above 4,000 feet along the Kaupo Trail and at Ulupalakua. The genus of some 200 species is widespread; it has many species native to Hawaii. Some kinds are well-known garden plants. Pittosporum tobira (Thunb.) Ait., a native of China and Japan, is a favorite shrub in California.

HAWAIIAN HAWTHORN, ’ULEI, Osteomeles anthyllidifolia Smith (Lindl). A spreading shrub with compound leaves and fragrant, small, white flowers, like apple blossoms, that may appear throughout the year. The fruit is white and contains five stony seeds. Plants growing on ash flats are very small. The strong but pliable wood of ulei was used for digging sticks, fish spears, and hoops to keep the mouths of fishing nets open.

HAWAIIAN RASPBERRY, ’AKALA, Rubus hawaiiensis A. Gray. Fig. 5. A shrub with attractive pink flowers abundant at Paliku. It is found also in Koolau Gap at the foot of Leleiwi Pali and elsewhere. The fruit, agreeable but somewhat bitter, is remarkable for its large size. It ripens about the Fourth of July or later. A trailing native, R. macraei A. Gray, sprawls in foggy Koolau Gap. Its large, dark fruits are bitter.

MAMANE, Sophora chrysophylla Salisb. Fig. 4. A common native shrub or small tree of the Bean Family both inside and outside the crater up to tree line. It is recognized by more or less downy, narrow, compound leaves, racemes of yellow flowers, and twisted pods that have four wings and are constricted between the seeds. Goats eat it greedily and quickly exterminate it in an area. The hard, durable wood was used by Hawaiians in many ways; today it is a principal firewood for the crater cabins. A tree in full bloom is a beautiful object. The height of the flowering season is mid-spring.

GERANIUM, NOHOANU, HINAHINA. Fig. 6, 7. Neurophyllodes tridens (Hillebr.) Degener & Greenwell is common above the park entrance and on the south wall within the crater. It sometimes becomes three feet tall and is readily identified by its silvery leaves, each of which has three small teeth on the end. The white flowers have purplish veins especially toward their centers. The blooming season is July to October. The silvery aspect of the plant, like the silversword, is imparted by a mesh of fine white hair that reflects the light of a passing car as effectively as the glass-beaded paint of directional signs.

Two native geranium relatives grow in Koolau Gap, N. ovatifolium (A. Gray) Degener & Greenwell and its variety superbum. The latter is common on the trail to Waikau not far from the foot of Leleiwi Pali.

One of the common plants growing on the crater floor is the exotic pink-flowered G. carolinianum var. australi (Benth.) Fosberg whose pointed fruiting bodies give it the common name, cranesbill.

HAWAIIAN HOLLY, KAWA’U, Ilex anomala Hook and Arn. The forest growing on the talus behind the Paliku cabins is the finest within the crater. A striking tree of this association is the Hawaiian holly that has dark, shiny, oval leaves with conspicuous networks of slightly depressed veins that make identification easy. Dense panicles of small, white flowers are followed by shiny, black drupes, like Christmas holly “berries.” The genus is that of the English holly. Curiously, the scientific name has been locally corrupted to ileck.

OLOMEA, Perrottetia sandwicensis A. Gray. A native shrub or small tree belonging to the same family as the bittersweet of the continent. The numerous tiny, round, red fruits, borne in panicles, suggest the relationship. The bright red venation of leaves and petioles make the plant easily recognized. I know of no plants within the park, but found many trees a thousand feet below the park boundary in Keanae Valley.

A’AL’II, Dodonaea eriocarpa Smith. Fig. 8. This is a common shrub in several varieties along the highway and in the western end of the crater. At Paliku and Kaupo Gap it becomes a tree up to 20 feet high and 8 inches in diameter. Its flowers are inconspicuous but clusters of dry, reddish fruit-capsules contrast, flower-like, with the surrounding green foliage. The fruits, abundant from July to September, are used for leis and dry bouquets. The hard brown heartwood was used for spears, pololu, daggers, pahoa, and other implements.

BEGONIA—PUAMAKANUI, Hillebrandia sandwicensis Oliv. (See illustration p. 53.) The only native begonia, found in wet ravines often by waterfalls. It grows profusely at Koolau a mile below Holua Cabin at the foot of the rain-drenched pali. This succulent herb has a tuberous rhizome, unbranched, slender stems, and hairy, toothed leaves 4-10 inches in diameter. From June to August it bears sprays of bright, pink and white flowers. This is one of the floral treasures of Hawaii.

TARWEED. Cuphea carthagenensis (Jacq.) McBride. A low, sticky, hairy perennial from tropical America widely spread at lower elevations in the park. It has red or green branches, small ovate leaves, and tiny but not unattractive pink flowers ¼ inch across. Plants in rock crevices on cliffs are tiny; on the crater floor, they may become a foot high and form a dense shrubby mat over a sizeable area. The plant belongs to the LYTHRACEAE or Crepe Myrtle Family.

’OHI’A LEHUA, Metrosideros collina (Forst.) A. Gray. Fig. 9. This, the commonest tree in the islands, consists of a swarm of hybrids of which the parentage is still unknown. It is scattered within the park to tree line. It is abundant in the eastern end of the crater and at Paliku. The beautiful flowers, mostly red, a few yellow, may appear throughout the year. Here spring seems to be the best season for them.

EVENING PRIMROSE, Raimannia odorata (Jacq.) Sprague & Riley. A slender, erect, hairy South American herb introduced forty years ago. It is a prolific bloomer, is widespread, and, in blooming season, the most conspicuous flower both at Park Headquarters and within the crater. The large, sulphur-yellow flowers appear at night, but wilt within the following day, turning reddish as they do so.

’APE’APE—Gunnera petaloidea Gaud. (See illustration p. 54.) A huge-leaved forest perennial with a massive prostrate stem which stands erect 3 or 4 feet at the tip. In the center of the leaves a tall stalk rises that bears hundreds of small yellow-brown flowers. I know of no plants within the crater, but some grow within a mile of the park boundary below Koolau Gap, as well as in adjacent wet valleys.

OLAPA, Cheirodendron trigynum (Gaud.) Heller. Fig. 10. This tree occurs in several varieties. The variety oblongum Sherff grows 30 feet tall around the cabins at Paliku, also at Ulupalakua. The variety mauiense Levl. common at Olinda, also grows in Kaupo Gap. The genus is Hawaiian but has a lone representative in the Marquesas. It is widespread in deep soils in all of the islands. The leaves are compound with 3 or 5 leaflets which, at Paliku, have reddish petioles. The panicles of small, green flowers are followed by black drupes. The leaves, bark, and fruit are said to have yielded a blue dye for staining tapa.

’OHELO, Vaccinium reticulatum Smith. Fig. 11. Hikers in the crater are grateful for the widespread ohelo bushes that yield pleasant fruits to be nibbled along the way. In areas rich in moisture, like Koolau Gap, large, maroon bell-shaped flowers droop from the axels of the leaves. The berries found in the eastern end of the crater appear to be largest. The bearing season appears at its height in mid-autumn. The plants along the Halemauu Trail constitute a distinct species, V. berberidifolium Skottsb.

PUKIAWE, Styphelia tameiameiae (Cham.) F. Muell. Fig. 12. A most abundant shrub, both inside and outside the crater, but near the top it is replaced by a trailing shrub that has been classified as S. douglasii (A. Gray) Hochr. The berries of this plant are white, pink, red, or mahogany brown; they are most abundant in winter, but some may be seen on the shrub throughout the year.

KOLEA, Suttonia lessertiana (A.DC.) Mez; syn. Rapanea lessertiana (A.DC.) Degener and Hosaka. Fig. 13. This variable tree grows up to 50 feet tall as one of the common trees around Paliku cabins. The thick leaves with short petioles, crowded near the ends of thick branches, have a beautiful roseate hue when young. The branches are studded with spurs on which grow small 5-parted flowers in clusters of three or more that are followed by dark, purplish-red or black fruits up to ¼ inch in diameter, often so numerous that the branch is completely hidden. Hawaiians made a red dye for tapa from the sap and bark. The crimson sap bleeds freely from a cut made deeply into the bark of a living tree.

SELFHEAL, Prunella vulgaris L., a common weed of Eurasian origin, is widespread in rocky, scrub cover, and at Paliku. A dense cluster of small, lipped, blue to purple flowers appears on the end of each upright stem.

PUA’AINAKA, Stenogyne rotundifolia A. Gray. This endemic long-branched shrub of the Mint Family is found only on Haleakala. Within the crater and on upper slopes it is trailing and is relatively rare, being most abundant in Kaupo Gap. The attractive, pale purple flowers in whorls of six are 1½ inches long and are covered with silky, white hair. They appear in late summer. Outside the crater, S. haliakalae Wawra is abundant in forests as a large, diffuse shrub that often forms a dense mat over surrounding shrubs. Another mint, Stenogyne crenata A. Gray, was collected by Skottsberg among shrubs on the south wall of the crater.

GROUNDCHERRY, Cape Gooseberry, POHA, Physalis peruviana L. A South American perennial herb, widely scattered throughout Hawaii, well-known for its round, orange, many-seeded, husk-enclosed fruits that are edible raw or preserved. It grows extensively in Kaupo Gap. The large yellow flowers with brownish spots near the center appear from June to late fall in this area.

PLANTAIN, LAUKAHI, Plantago sp. Plantain is a hardy, cosmopolitan, stemless weed forming a rosette of broadly oval leaves, 1-10 inches long, near the ground. The tiny flowers and seed capsules are borne as cylindrical heads at the ends of tall stalks. The plant is widespread along the side of the road as well as within the crater. Of several hundred species of Plantago, four or more are endemic to Hawaii. One of these, P. princeps Cham. & Schl., is a shrub several feet high with tufts of long narrow leaves at the ends of the branches. It grows on cliffs at Kaupo Gap. Other plantains with thick leaves, silky underneath, creep on the ground in Koolau Gap.

KUKAENENE, Coprosma ernodioides A. Gray. Fig. 15. A common woody shrub with long trailing branches that send up short, erect, densely foliose branchlets at each node. The awl-shaped leaves are rigid and dark-green. The fruits are shiny black drupes which are a favorite food of the native goose, nene.

PILO, Coprosma montana Hillebr. Fig. 14. One of the commonest shrubs throughout the crater and from Park Headquarters to 9,000 feet outside the crater. It is a small tree up to 20 feet tall in Kaupo Gap. As a shrub the ascending tips look like jets shot up from densely foliose branchlets. The alternate, small, thick leaves have conspicuous nerves impressed on the upper face. Below each pair of leaves is a pair of triangular bracts, stipules, with cilia on the upper border. The greenish, inconspicuous flowers are followed by showy, bright orange, yellow, or red fruits which make the plant a subject attractive to color photographers in fall.

MANONO, Gouldia terminalis (H. & A.) Hillebr. A shrub or small tree growing on the talus above the Paliku cabins. It has shiny, opposite leaves and dense terminal clusters of greenish, four-lobed, cup-shaped flowers that are followed by small black berries. It blooms in late summer. The genus is one of three in the Coffee Family that are endemic to Hawaii.

CATCHFLY, Silene struthioloides A. Gray. Fig. 16. A plant that is typical only of arid Cinders and ash on East Maui and on the island of Hawaii. With the silversword as its companion in Haleakala Crater the plants are found at the bases of barren cones. They show neat adaptation to their stark home. The Haleakala plants, as illustrated, are low and compact, but those growing at Kilauea Crater bear only a few awl-shaped leaves and resemble dead twigs. The thick tap roots are sweet and edible. About 250 species belong to this genus, a member of the Pink or Carnation Family. A well-known introduced weed, the English catchfly, Silene anglica L., was reported by Degener at 10,000 feet on Haleakala.

’OHA, Labelia grayana E. Wimm. A low plant with woody trailing stems with knobby leaf scars and ending with a crowded arrangement of silvery, linear leaves, 4-8 inches long and crowned with densely flowered racemes, 6-15 inches long. The flowers are lilac-blue with a satiny sheen. The plant is not uncommon on wet pali from 5,000-7,000 feet at Paliku, Kaupo and Koolau Gaps, and in the northwestern end of the crater. It is a glorious plant worth hunting for and going miles to see.

NAUPAKA. Scaevola chamissoniana Gaud. This shrub was noted only on the east side of Kaupo Gap. This is a varying species found up to the 6,000-foot elevation. It is not common and blooms in summer. The white flowers with purplish streaks are slit to the base on the upper side. They look like flower-halves rather than complete corollas. There are several legends about the peculiar flower, each dealing with lovers separated from each other. In a song composed about it, the lovers were forceably parted, so the girl divided a perfect corolla, giving one half to her lover while keeping the other half herself. One of the lovers carried the flower to the sea, naupaka kahakai, the other to the mountain, naupaka kuahiwi, where the plants are found today.

MAUI WORMWOOD, Artemisia mauiensis (A. Gray) Skottsb. Fig. 17. Typical of Maui and found only on Haleakala, this hoary ornamental shrub, usually 2-3 feet high, perches on cliffs usually above the reach of man. It has a densely-branched crown with silvery leaves that are aromatic and bitter. The leaves are composed of thin segments that are covered with a mat of cottony hair, giving the plant a silvery appearance. The small orange flowers are borne in terminal panicles.

The Hawaiians call the wormwood AHINAHINA, applied also to silversword, geranium, and other gray plants. The basic word refers to the color of silvery-gray hair, the connection being obvious. Hawaiians use the pounded leaves to relieve asthma. The genus is large, having some 250 species that are generally found in arid regions. The sagebrush of the western states, A. tridentata Nutt., is the best known to most park visitors.

KO’OKO’OLAU, Bidens sp. Like Artemisia, the genus is a huge one with over 200 species and belongs to the Composite Family which includes dandelions, daisies, and sunflowers. E. E. Sherff of the Chicago Museum of Natural History, a specialist on the genus, lists sixty species native to Hawaii. Native KO’OKO’OLAU are shrubby and often of great beauty. This is true of B. campylotheca pentamera Sherff which sprawls over the vegetation in Koolau Gap. It has fern-like leaves and large, pretty, yellowish flower-heads. Hawaiians use the tips of young plants for tea, often in preference over imported tea.

Besides the native varieties, three introduced species grow in the islands, including beggar ticks or Spanish needles, B. pilosa L. It is a nuisance, as the three-pronged fruits that give it the common name readily attach themselves to clothing as well as to fur of passing animals.

KUPAOA, NA’ENA’E. Fig. 18, 19. Several kinds of composite shrubs are called by these names, both of which mean fragrance or perfume. They were used for scenting tapa. They belong to the endemic genera Dubautia and Railliardia, both of which have species found in the park. Dubautia plantagiena var. platyphylla Hillebr. Gaud., a shrub at 6,000 feet in Kaupo and Koolau Gaps, has linear leaves 4-8 inches long, with 7-13 conspicuous nerves. It is a small tree below Nianiau Crater. It is a handsome sight when in flower; the flower-heads are yellow. In general appearance it is much like that of Railliardia platyphylla A. Gray which grows in cinders and ash, mostly inside the crater. At 8,000 feet, R. platyphylla becomes a straggling shrub. The commonest member of this group, Railliardia menziesii A. Gray, is a shrub in the crater and on the rim from 8,000-10,000 feet. Between 6,000 and 7,000 feet it is a tree up to 20 feet tall. The dark green, pointed, linear, fleshy leaves are ranked in vertical rows of four on upright stems and branches. Its dark yellow flower-heads are borne in panicles. R. scabra DC., found in Kaupo Gap and on Leleiwi Pali, does not have the regular leaf arrangement.

PAMAKANI-HAOLE, Eupatorium glandulosum HBK. Eupatorium is another huge genus of the Composite Family with several hundred species, mostly from tropical America. Five species have been introduced into Hawaii of which two are bad pests. E. glandulosum, a native of Mexico, spread rapidly on Maui, crowding out desirable plants and making pasturelands worthless. A parasitic insect, Procecidochares utilis Stone, was introduced in 1944 to combat it. This insect belongs to the order DIPTERA, the flies, gnats, midges, and mosquitoes, and to the family TRYPETIDAE, that includes the Mediterranean fruitfly and the common “apple-worm” which is actually the larva of a fly. The trypetids infest living plants, frequently causing galls, and have piercing ovipositors, often prominent, with which females deposit eggs beneath the skin of their host.

It is not expected that the studied introduction of a parasite will result in the extermination of a host, but rather that it will check unbridled increase and spread. The method is termed “biological control.” It has been successfully adopted against several menaces, such as cactus, lantana, and a fern-weevil, Syragrius fulvitarsis Pascoe, that kills amaumau ferns in the Kilauea Crater area.

Pamakani abound throughout the crater, even clinging as tiny starvelings in small cracks in cliffs, like those along Halemauu Trail. Every plant shows swellings in which the little maggots live, and exit holes through which the new adults emerge. Many plants bear only few leaves and fight tenaciously for survival. Big plants may be seen along Kaupo Trail. Pamakani means wind-blown, in reference to the method of seed dispersal.

HAIRY CAT’S-EAR, “Dandelion,” Hypochoeris radicata L. A common composite, native to the Mediterranean, found in abundance throughout the park. Its narrow leaves with yellow hairs form flat rosettes. A branching, leafless stalk up to a foot or more tall bears yellow flower-heads, an inch in diameter, that resemble the well-known dandelion, Taraxacum officinale Weber, by which common name many call the cat’s-ear. The hairless, H. glabra L. is a smaller plant with smooth leaves and flower-heads ¼ inch in diameter. The cat’s-ear is a favorite food of the Hawaiian goose, nene. Gosmer is a common name used locally.

WOOD GROUNDSEL, Senecio sylvaticus L., a native of Europe, is a branching, weedy herb, 1-2 feet high, abundant along the foot of Kalahaku Pali. It has irregularly lobed leaves and small yellow flowers in a tight flower head, ? inch long and ½ inch in diameter.

TETRAMALOPIUM. Fig. 20, illustration p. 53. This endemic genus has a dozen species, two of which are a pride of Haleakala. The small leaves are narrow and crowded at branch ends. The showy flowerheads of white, pinkish, or lavender ray florets surround a disk of purplish central florets. T. humile (A. Gray) Hillebr. is found inside and outside the crater between 6,000-9,000 feet. It is a small, low, shrubby plant with narrow, spoon-shaped leaves quite covered with sticky, curly hair. The plant growing in cracks between rocks at the very summit is stiffer and considerably different.

SILVERSWORD, AHINAHINA, Argyroxiphium sandwicense DC. Illustrations on cover and pages iv, 47. As famous as the crater itself, and almost as well-known, the silversword is regarded as typical of Haleakala, although its natural range embraces Mauna Loa, Mauna Kea, and Hualalai from 7,000 to 12,000 feet. A lustrous silvery down thickly covers all leaves and makes the plant exceedingly beautiful. It is highly evolved to withstand the extreme dryness of cinder cones and the intense sunlight of lofty elevation. Surprising to most visitors on first acquaintance is the fact that it bears no relationship to the yucca of the Lily Family, but belongs to the COMPOSITAE along with sunflowers, asters, and chrysanthemums. Like Dubautia and Railliardia, it has no close relatives outside of the islands. Wilkesia, endemic to Kauai, is so closely similar that some authorities class it in the genus Argyroxphium.

The Haleakala silversword has a short, simple, woody stem 2-3 inches in diameter crowded with thick, dagger-like leaves arranged spirally around it. After growth from 7 to as much as 20 years, a foliose raceme 3-8 feet high develops on which 100-500 flower-heads nod. Each has a central disk of hundreds of bright yellow florets surrounded by a score of short reddish-purple ray florets. The flowering season is from June through October. The whole plant dies after flowering but once.

The greensword, A. virescens Hillebr., a much rarer plant with green leaves, once grew from 6,000 to 9,000 feet. Today it has all but vanished; a few plants are still to be found in Koolau Gap just outside the park, and between rocks on the edges of cliffs at Kaupo Gap.

Visitors once gathered specimens of silversword as evidence that they had been up the mountain, even as the mountaineer gathers edelweiss in Switzerland. Thoughtless people uprooted the silvery globes merely to watch them tumble down the slopes. Goats eat the growing heart of the plant. Grazing stock are incompatible with its existence. It has many insect parasites (see The Insect Life). All of these, singly and in combination, threaten the existence of the species.

The outer slopes of the mountain had thousands of silverswords according to early accounts, while today they are rare. In the crater, some of the cones were so thickly covered that they appeared to be bathed in moonlight. Except for strong, enlightened action, this beautiful plant might have completely vanished from the mountain.

                                                                                                                                                                                                                                                                                                           

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