Natural Selection.

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We have for the moment provisionally admitted the theory of natural selection, in order to see [pg 155] whether it could be included in a religious interpretation of things. But in reality such an admission is not to be thought of, in face of what is at present so apparent—the breaking down of this hypothesis, which has been upheld with so much persistence. We shall have to occupy ourselves with this later on. In the meantime a few more remarks must be added to what has been already said.

It might be said, paradoxically, that the worst fate that could befall this hypothesis would be to be proved, for then it would be most certainly refuted. What we mean is this: If it is really “utility” that rules the world and things, there can be no certainty and objectivity of knowledge, no guarantee of truth. The “struggle for existence” is not concerned with selecting beings who see the world as it is. It selects only the interpretation and conception of the environment that is most serviceable for the existence and maintenance of the species. But there is nothing to guarantee that the “true” knowledge will also be the most useful. It might quite well be that an entirely subjective and in itself wholly false interpretation would be the most serviceable. And if, by some extraordinary chance, the selected interpretation should be also the true one, there would be no means of establishing the fact. And what is true of this interpretation is true also of all theories that are derived from it, for example of the theory of selection itself.

Furthermore, a great part, perhaps the greatest part [pg 156] of the confidence placed in the theory of selection is due to an involuntary, but entirely fallacious habit of crediting it with the probabilities in favour of the doctrine of descent. The main arguments in favour of evolution and descent are very often, though unwittingly, adduced in support of Darwinism in particular. This is a great mistake. Take, for instance, the evidence of the “palÆontological” record. It affords hundreds of proofs of evolution, but not a single proof of selection. Its “intermediate” and “connecting links” do possibly prove the affiliation of species and the validity of genealogical trees. But precisely the “intermediate links” which selection requires—the myriads of forms of life which were not successfully adapted, the unfit competitors in the struggle for existence which must have accompanied the favourably adapted variants from step to step, from generation to generation—these are altogether awanting.

Another circumstance seems to us to have been entirely overlooked, and it is one which gives the theory of selection an inevitable appearance of truth, even if it is essentially false, and thus makes it very difficult to refute. Assuming that the recognition of teleological factors is valid, that there is an inward law of development, that “Moses” or whoever one will was undoubtedly right, it is self-evident that, because of the indubitable over-production of organisms, there would even then be a struggle for existence on an immense scale, and that it would have a far-reaching [pg 157] “selective” influence, because of the relative plasticity of many forms of life. Beyond doubt it would, in the course of Æons, have applied its shears to many forms of life, and probably there would be no organisms, organs, or associations in the evolution of the ultimate form of which it had not energetically co-operated. Its influence would, perhaps, be omnipresent, yet it might be far from being the all-sufficient factor in evolution; indeed, as far as the actual impulse of evolution is concerned, it might be a mere accessory. Unless we are to think of the forms of life as wholly passive and wooden, the struggle for existence must necessarily be operative, and the magnitude of its results, and their striking and often bizarre outcome, will tend ever anew to conceal the fact that the struggle is after all only an inevitable accompaniment of evolution. And thus we understand how it is that interpretations from the point of view of an inward law of development, of orthogenesis, or of teleology, notwithstanding their inherent validity, have À priori always had a relatively difficult position as compared with the Darwinian view.

It is usual to speak of the “all-sufficiency of natural selection,” yet the champion of the selection-theory admits, as he needs must, that the struggle for existence and selection can of themselves create absolutely nothing, no new character, no new or higher combination of the vital elements; they can only take what is already given; they can only select and eliminate [pg 158] among the wealth of what is offered.38 And the offerer is Life itself by virtue of its mysterious capacity for boundless and inexhaustible variability, self-enrichment and increase. The “struggle for existence” only digs the bed through which life's stream flows, draws the guiding-line, and continually stimulates it to some fresh revelation of its wealth. But this wealth was there from the beginning; it was, to use the old word, “potential” in the living, and included with it in the universal being from which life was called forth. The struggle for existence is only the steel which strikes the spark from the flint; is, with its infinite forms and components, only the incredibly complex channel through which life forces its way upwards. If we keep this clearly in mind, the alarming and ominous element in the theory shrinks to half its dimensions.

And, finally, if we can rid ourselves of the peculiar fascination which this theory exercises, we soon begin to discover what extraordinary improbability and fundamental artificiality it implies. “Utility” is maintained to be that which absolutely, almost tyrannically, determines form and development in the realm of the living. Is this an idea that finds any analogy elsewhere in nature? Those who uphold the theory most strongly are wont to compare the development of organisms to crystal-formation in order in some way to tack on the living to the not-living. Crystal-formation, [pg 159] with its processes of movement and form-development, is, they say, a kind of connecting link between the living and the not-living. And in truth we find here, as in the realm of life, species-formation, development into individuals, stages and systems. But all this takes place without any hint of “struggle for existence,” of laboriously “selective” processes, or of ingenious accumulation of “variations.” The “species” of crystals are formed not according to utility, but according to inherent, determining laws of development, to which the diversity of their individual appearances is due. If “Life” were only a higher potential of what is already stirring in crystallisation, as this view suggests, then we should expect to find fixed tendencies, determined from within, in accordance with which life would pass through the cycle of its forms and possibilities, and rise spontaneously through gradual stages.

[pg 160]

Let us turn now to the other side. What is opposed to Darwinism in the biological investigations of the experts of to-day is in part simple criticism of the Darwinian position as a whole or in some of its details, and in part constructive individual theories and interpretations of the evolution of organisms.

A. Fleischmann's book, “Die Darwinsche Theorie,”39 is professedly only critical. He suggests no theory of his own as to the evolution of life in contrast to Darwin's; for, as we have already seen in connection with his earlier book, “Die Deszendenztheorie,” he denies evolution altogether. His agnostic position is maintained, if possible, more resolutely than before. Natural science, according to him, must keep to facts. Drawing conclusions and spinning theories is inexact, and distracts from objective study. The Darwinian theory of selection seems to him a particularly good example of this, for it is built up À priori on theories [pg 161] and hypotheses, it stands apart from experimentation, and it twists facts forcibly to its own ends. It has, however, to be acknowledged that Fleischmann's book is without any “apologetic” intentions. It holds equally aloof from teleology. To seek for purposes and aims in nature he holds to be outside the business of science, as Kant's “Critique of Judgment” suffices to show. After having been more than a decade under the charm of the theory of selection, Fleischmann knows its fascination well, but he now regards it as so erroneous that no one who wishes to do serious work should concern himself about it at all. Point by point he follows all the details of Darwin's work, and seeks to analyse the separate views and theories which go to make up Darwinism as a whole. Darwin's main example of the evolution of the modern races of pigeons from one ancestral form, Columba livia, is, according to Fleischmann, not only unproved but unprovable.40 For this itself is not a unified type. The process of “unconscious selection” by man is obscure, and it is not demonstrable, especially in regard to pigeon-breeding. It is a hazy idea which cannot be transferred to the realm of nature. The Malthusian assumption of the necessity of the struggle for existence is erroneous. Malthus was wrong in his law of population as applied [pg 162] to human life, and Darwin was still more mistaken when he transferred it to the organic world in general. It was mere theory. Statistics should have been collected, and observations instead of theories should have been sought for. The alleged superabundance of organisms is not a fact. The marvellously intertwined conditions in the economy of nature make the proportion of supply and demand relatively constant. And even when there is actual struggle for existence, advantages of situation,41 which are quite indifferent as far as selection is concerned, are much more decisive than any variational differences. The theory does not explain the first origin of new characters, which can only become advantageous when they have attained to a certain degree of development. As to the illustrations of the influence of selection given by Darwin, from the much discussed fictitious cases, in which the fleet stags select the lithe wolf, to the marvellous mutual adaptations of insects and flowers, Fleischmann objects that there is not even theoretical justification for any one of them. The spade-like foot of the mole is not “more useful” than the form of foot which probably preceded it (cf. Goette), it is merely “different.” For when the mole took to burrowing in the earth and adapting itself to that mode of life, it ipso facto forfeited all the advantages of living above ground. The postulated myriads of less well-adapted forms of life are no more to be found to-day than they are in the fauna and flora of [pg 163] palÆontological times. The famous giraffe story has already been disposed of by Mivart's objections. As to the whales, it is objected that the earliest stages of their whalebone and their exaggerated nakedness can have been of no use, and a series of other alleged selective effects of “utility” are critically analysed. The refutation of the most brilliant chapter in the Darwinian theory, that on protective coloration and mimicry, is very insufficient. A long concluding chapter sums up the fundamental defects of the Darwinian theory.

For the most part, Fleischmann simply brings forward objections which have been urged against the theory of selection from the first, either by naturalists or from other quarters. The chief and the most fatal of these which are still current are the following: The theory of selection does not explain the actually existing discontinuity of species. The real characteristics which distinguish species from species are in innumerable cases quite indifferent from the point of view of “utility” (NÄgeli, Bateson). “Selection preserves the good and weeds out the bad.” But where does the good come from? (De Vries). The first beginnings of what may later be useful are almost always useless. The theory of selection might perhaps explain the useful qualities, but not the superfluous, useless, or directly injurious characters which actually exist. Confirmation of the theory of descent may be found in the palÆontological record, but it affords none of the theory of selection. Natural selection is continually being neutralised by [pg 164] subsequent inter-crossing and reversion. Natural selection may indeed prevent degeneration within the limits of the species by weeding out what is weak and bad, but it is powerless beyond these limits, and so forth.42

These ever-repeated and ever-increasing objections are purely critical. As this is true of Fleischmann's whole book, it is therefore unsatisfactory. It leaves everything in the mist, and puts nothing in place of what it attempts to demolish. But attempts are being made in other quarters, especially among the Lamarckians, to build up an opposition theory.


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