THE FIBRES OF CORTI.

Previous

Whoever has roamed through a beautiful country knows that the tourist's delights increase with his progress. How pretty that wooded dell must look from yonder hill! Whither does that clear brook flow, that hides itself in yonder sedge? If I only knew how the landscape looked behind that mountain! Thus even the child thinks in his first rambles. It is also true of the natural philosopher.

The first questions are forced upon the attention of the inquirer by practical considerations; the subsequent ones are not. An irresistible attraction draws him to these; a nobler interest which far transcends the mere needs of life. Let us look at a special case.

For a long time the structure of the organ of hearing has actively engaged the attention of anatomists. A considerable number of brilliant discoveries has been brought to light by their labors, and a splendid array of facts and truths established. But with these facts a host of new enigmas has been presented.

Whilst in the theory of the organisation and functions of the eye comparative clearness has been attained; whilst, hand in hand with this, ophthalmology has reached a degree of perfection which the preceding century could hardly have dreamed of, and by the help of the ophthalmoscope the observing physician penetrates into the profoundest recesses of the eye, the theory of the ear is still much shrouded in mysterious darkness, full of attraction for the investigator.

Look at this model of the ear. Even at that familiar part by whose extent we measure the quantity of people's intelligence, even at the external ear, the problems begin. You see here a succession of helixes or spiral windings, at times very pretty, whose significance we cannot accurately state, yet for which there must certainly be some reason.

Fig. 6.

The shell or concha of the ear, a in the annexed diagram, conducts the sound into the curved auditory passage b, which is terminated by a thin membrane, the so-called tympanic membrane, e. This membrane is set in motion by the sound, and in its turn sets in motion a series of little bones of very peculiar formation, c. At the end of all is the labyrinth d. The labyrinth consists of a group of cavities filled with a liquid, in which the innumerable fibres of the nerve of hearing are imbedded. By the vibration of the chain of bones c, the liquid of the labyrinth is shaken, and the auditory nerve excited. Here the process of hearing begins. So much is certain. But the details of the process are one and all unanswered questions.

To these old puzzles, the Marchese Corti, as late as 1851, added a new enigma. And, strange to say, it is this last enigma, which, perhaps, has first received its correct solution. This will be the subject of our remarks to-day.

Corti found in the cochlea, or snail-shell of the labyrinth, a large number of microscopic fibres placed side by side in geometrically graduated order. According to KÖlliker their number is three thousand. They were also the subject of investigation at the hands of Max Schultze and Deiters.

A description of the details of this organ would only weary you, besides not rendering the matter much clearer. I prefer, therefore, to state briefly what in the opinion of prominent investigators like Helmholtz and Fechner is the peculiar function of Corti's fibres. The cochlea, it seems, contains a large number of elastic fibres of graduated lengths (Fig. 7), to which the branches of the auditory nerve are attached. These fibres, called the fibres, pillars, or rods of Corti, being of unequal length, must also be of unequal elasticity, and, consequently, pitched to different notes. The cochlea, therefore, is a species of pianoforte.

Fig. 7.

What, now, may be the office of this structure, which is found in no other organ of sense? May it not be connected with some special property of the ear? It is quite probable; for the ear possesses a very similar power. You know that it is possible to follow the individual voices of a symphony. Indeed, the feat is possible even in a fugue of Bach, where it is certainly no inconsiderable achievement. The ear can pick out the single constituent tonal parts, not only of a harmony, but of the wildest clash of music imaginable. The musical ear analyses every agglomeration of tones.

The eye does not possess this ability. Who, for example, could tell from the mere sight of white, without a previous experimental knowledge of the fact, that white is composed of a mixture of other colors? Could it be, now, that these two facts, the property of the ear just mentioned, and the structure discovered by Corti, are really connected? It is very probable. The enigma is solved if we assume that every note of definite pitch has its special string in this pianoforte of Corti, and, therefore, its special branch of the auditory nerve attached to that string. But before I can make this point perfectly plain to you, I must ask you to follow me a few steps into the dry domain of physics.

Look at this pendulum. Forced from its position of equilibrium by an impulse, it begins to swing with a definite time of oscillation, dependent upon its length. Longer pendulums swing more slowly, shorter ones more quickly. We will suppose our pendulum to execute one to-and-fro movement in a second.

This pendulum, now, can be thrown into violent vibration in two ways; either by a single heavy impulse, or by a number of properly communicated slight impulses. For example, we impart to the pendulum, while at rest in its position of equilibrium, a very slight impulse. It will execute a very small vibration. As it passes a third time its position of equilibrium, a second having elapsed, we impart to it again a slight shock, in the same direction with the first. Again after the lapse of a second, on its fifth passage through the position of equilibrium, we strike it again in the same manner; and so continue. You see, by this process the shocks imparted augment continually the motion of the pendulum. After each slight impulse, the pendulum reaches out a little further in its swing, and finally acquires a considerable motion.[8]

But this is not the case under all circumstances. It is possible only when the impulses imparted synchronise with the swings of the pendulum. If we should communicate the second impulse at the end of half a second and in the same direction with the first impulse, its effects would counteract the motion of the pendulum. It is easily seen that our little impulses help the motion of the pendulum more and more, according as their time accords with the time of the pendulum. If we strike the pendulum in any other time than in that of its vibration, in some instances, it is true, we shall augment its vibration, but in others again, we shall obstruct it. Our impulses will be less effective the more the motion of our own hand departs from the motion of the pendulum.

What is true of the pendulum holds true of every vibrating body. A tuning-fork when it sounds, also vibrates. It vibrates more rapidly when its sound is higher; more slowly when it is deeper. The standard A of our musical scale is produced by about four hundred and fifty vibrations in a second.

I place by the side of each other on this table two tuning-forks, exactly alike, resting on resonant cases. I strike the first one a sharp blow, so that it emits a loud note, and immediately grasp it again with my hand to quench its note. Nevertheless, you still hear the note distinctly sounded, and by feeling it you may convince yourselves that the other fork which was not struck now vibrates.

I now attach a small bit of wax to one of the forks. It is thrown thus out of tune; its note is made a little deeper. I now repeat the same experiment with the two forks, now of unequal pitch, by striking one of them and again grasping it with my hand; but in the present case the note ceases the very instant I touch the fork.

What has happened here in these two experiments? Simply this. The vibrating fork imparts to the air and to the table four hundred and fifty shocks a second, which are carried over to the other fork. If the other fork is pitched to the same note, that is to say, if it vibrates when struck in the same time with the first, then the shocks first emitted, no matter how slight they may be, are sufficient to throw the second fork into rapid sympathetic vibration. But when the time of vibration of the two forks is slightly different, this does not take place. We may strike as many forks as we will, the fork tuned to A is perfectly indifferent to their notes; is deaf, in fact, to all except its own; and if you strike three, or four, or five, or any number whatsoever, of forks all at the same time, so as to make the shocks which come from them ever so great, the A fork will not join in with their vibrations unless another fork A is found in the collection struck. It picks out, in other words, from all the notes sounded, that which accords with it.

The same is true of all bodies which can yield notes. Tumblers resound when a piano is played, on the striking of certain notes, and so do window panes. Nor is the phenomenon without analogy in other provinces. Take a dog that answers to the name "Nero." He lies under your table. You speak of Domitian, Vespasian, and Marcus Aurelius Antoninus, you call upon all the names of the Roman Emperors that occur to you, but the dog does not stir, although a slight tremor of his ear tells you of a faint response of his consciousness. But the moment you call "Nero" he jumps joyfully towards you. The tuning-fork is like your dog. It answers to the name A.

You smile, ladies. You shake your heads. The simile does not catch your fancy. But I have another, which is very near to you: and for punishment you shall hear it. You, too, are like tuning-forks. Many are the hearts that throb with ardor for you, of which you take no notice, but are cold. Yet what does it profit you! Soon the heart will come that beats in just the proper rhythm, and then your knell, too, has struck. Then your heart, too, will beat in unison, whether you will or no.

The law of sympathetic vibration, here propounded for sounding bodies, suffers some modification for bodies incompetent to yield notes. Bodies of this kind vibrate to almost every note. A high silk hat, we know, will not sound; but if you will hold your hat in your hand when attending your next concert you will not only hear the pieces played, but also feel them with your fingers. It is exactly so with men. People who are themselves able to give tone to their surroundings, bother little about the prattle of others. But the person without character tarries everywhere: in the temperance hall, and at the bar of the public-house—everywhere where a committee is formed. The high silk hat is among bells what the weakling is among men of conviction.

A sonorous body, therefore, always sounds when its special note, either alone or in company with others, is struck. We may now go a step further. What will be the behaviour of a group of sonorous bodies which in the pitch of their notes form a scale? Let us picture to ourselves, for example (Fig. 8), a series of rods or strings pitched to the notes c d e f g.... On a musical instrument the accord c e g is struck. Every one of the rods of Fig. 8 will see if its special note is contained in the accord, and if it finds it, it will respond. The rod c will give at once the note c, the rod e the note e, the rod g the note g. All the other rods will remain at rest, will not sound.

We need not look about us long for such an instrument. Every piano is an instrument of this kind, with which the experiment mentioned may be executed with splendid success. Two pianos stand here by the side of each other, both tuned alike. We will employ the first for exciting the notes, while we will allow the second to respond; after having first pressed upon the loud pedal, so as to render all the strings capable of motion.

Every harmony struck with vigor on the first piano is distinctly repeated on the second. To prove that it is the same strings that are sounded in both pianos, we repeat the experiment in a slightly changed form. We let go the loud pedal of the second piano and pressing on the keys c e g of that instrument vigorously strike the harmony c e g on the first piano. The harmony c e g is now also sounded on the second piano. But if we press only on one key g of one piano, while we strike c e g on the other, only g will be sounded on the second. It is thus always the like strings of the two pianos that excite each other.

The piano can reproduce any sound that is composed of its musical notes. It will reproduce, for example, very distinctly, a vowel sound that is sung into it. And in truth physics has proved that the vowels may be regarded as composed of simple musical notes.

You see that by the exciting of definite tones in the air quite definite motions are set up with mechanical necessity in the piano. The idea might be made use of for the performance of some pretty pieces of wizardry. Imagine a box in which is a stretched string of definite pitch. This is thrown into motion as often as its note is sung or whistled. Now it would not be a very difficult task for a skilful mechanic to so construct the box that the vibrating cord would close a galvanic circuit and open the lock. And it would not be a much more difficult task to construct a box which would open at the whistling of a certain melody. Sesame! and the bolts fall. Truly, we should have here a veritable puzzle-lock. Still another fragment rescued from that old kingdom of fables, of which our day has realised so much, that world of fairy-stories to which the latest contributions are Casselli's telegraph, by which one can write at a distance in one's own hand, and Prof. Elisha Gray's telautograph. What would the good old Herodotus have said to these things who even in Egypt shook his head at much that he saw? ??? ??e ?? p?sta, just as simple-heartedly as then, when he heard of the circumnavigation of Africa.

A new puzzle-lock! But why invent one? Are not we human beings ourselves puzzle-locks? Think of the stupendous groups of thoughts, feelings, and emotions that can be aroused in us by a word! Are there not moments in all our lives when a mere name drives the blood to our hearts? Who that has attended a large mass-meeting has not experienced what tremendous quantities of energy and motion can be evolved by the innocent words, "Liberty, Equality, Fraternity."

But let us return to the subject proper of our discourse. Let us look again at our piano, or what will do just as well, at some other contrivance of the same character. What does this instrument do? Plainly, it decomposes, it analyses every agglomeration of sounds set up in the air into its individual component parts, each tone being taken up by a different string; it performs a real spectral analysis of sound. A person completely deaf, with the help of a piano, simply by touching the strings or examining their vibrations with a microscope, might investigate the sonorous motion of the air, and pick out the separate tones excited in it.

The ear has the same capacity as this piano. The ear performs for the mind what the piano performs for a person who is deaf. The mind without the ear is deaf. But a deaf person, with the piano, does hear after a fashion, though much less vividly, and more clumsily, than with the ear. The ear, thus, also decomposes sound into its component tonal parts. I shall now not be deceived, I think, if I assume that you already have a presentiment of what the function of Corti's fibres is. We can make the matter very plain to ourselves. We will use the one piano for exciting the sounds, and we shall imagine the second one in the ear of the observer in the place of Corti's fibres, which is a model of such an instrument. To every string of the piano in the ear we will suppose a special fibre of the auditory nerve attached, so that this fibre and this alone, is irritated when the string is thrown into vibration. If we strike now an accord on the external piano, for every tone of that accord a definite string of the internal piano will sound and as many different nervous fibres will be irritated as there are notes in the accord. The simultaneous sense-impressions due to different notes can thus be preserved unmingled and be separated by the attention. It is the same as with the five fingers of the hand. With each finger I can touch something different. Now the ear has three thousand such fingers, and each one is designed for the touching of a different tone.[9] Our ear is a puzzle-lock of the kind mentioned. It opens at the magic melody of a sound. But it is a stupendously ingenious lock. Not only one tone, but every tone makes it open; but each one differently. To each tone it replies with a different sensation.

More than once it has happened in the history of science that a phenomenon predicted by theory, has not been brought within the range of actual observation until long afterwards. Leverrier predicted the existence and the place of the planet Neptune, but it was not until sometime later that Galle actually found the planet at the predicted spot. Hamilton unfolded theoretically the phenomenon of the so-called conical refraction of light, but it was reserved for Lloyd some time subsequently to observe the fact. The fortunes of Helmholtz's theory of Corti's fibres have been somewhat similar. This theory, too, received its substantial confirmation from the subsequent observations of V. Hensen. On the free surface of the bodies of Crustacea, connected with the auditory nerves, rows of little hairy filaments of varying lengths and thicknesses are found, which to some extent are the analogues of Corti's fibres. Hensen saw these hairs vibrate when sounds were excited, and when different notes were struck different hairs were set in vibration.

I have compared the work of the physical inquirer to the journey of the tourist. When the tourist ascends a new hill he obtains of the whole district a different view. When the inquirer has found the solution of one enigma, the solution of a host of others falls into his hands.

Surely you have often felt the strange impression experienced when in singing through the scale the octave is reached, and nearly the same sensation is produced as by the fundamental tone. The phenomenon finds its explanation in the view here laid down of the ear. And not only this phenomenon but all the laws of the theory of harmony may be grasped and verified from this point of view with a clearness before undreamt of. Unfortunately, I must content myself to-day with the simple indication of these beautiful prospects. Their consideration would lead us too far aside into the fields of other sciences.

The searcher of nature, too, must restrain himself in his path. He also is drawn along from one beauty to another as the tourist from dale to dale, and as circumstances generally draw men from one condition of life into others. It is not he so much that makes the quests, as that the quests are made of him. Yet let him profit by his time, and let not his glance rove aimlessly hither and thither. For soon the evening sun will shine, and ere he has caught a full glimpse of the wonders close by, a mighty hand will seize him and lead him away into a different world of puzzles.

Respected hearers, science once stood in an entirely different relation to poetry. The old Hindu mathematicians wrote their theorems in verses, and lotus-flowers, roses, and lilies, beautiful sceneries, lakes, and mountains figured in their problems.

"Thou goest forth on this lake in a boat. A lily juts forth, one palm above the water. A breeze bends it downwards, and it vanishes two palms from its previous spot beneath the surface. Quick, mathematician, tell me how deep is the lake!"

Thus spoke an ancient Hindu scholar. This poetry, and rightly, has disappeared from science, but from its dry leaves another poetry is wafted aloft which cannot be described to him who has never felt it. Whoever will fully enjoy this poetry must put his hand to the plough, must himself investigate. Therefore, enough of this! I shall reckon myself fortunate if you do not repent of this brief excursion into the flowered dale of physiology, and if you take with yourselves the belief that we can say of science what we say of poetry,

"Who the song would understand,
Needs must seek the song's own land;
Who the minstrel understand
Needs must seek the minstrel's land."
                                                                                                                                                                                                                                                                                                           

Clyx.com


Top of Page
Top of Page