General principles of classification of tertiary strata — Detached formations scattered over Europe — Strata of Paris and London — More modern groups — Peculiar difficulties in determining the chronology of tertiary formations — Increasing proportion of living species of shells in strata of newer origin — Terms Eocene, Miocene, and Pliocene — Post-Pliocene strata — Recent or human period — Older Post-Pliocene formations of Naples, Uddevalla, and Norway — Ancient upraised delta of the Mississippi — Loess of the Rhine. Before describing the most modern of the sets of strata enumerated in the tables given at the end of the last chapter, it will be necessary to say something generally of the mode of classifying the formations called tertiary. The name of tertiary has been given to them, because they are all posterior in date to the rocks termed "secondary," of which the chalk constitutes the newest group. These tertiary strata were at first confounded, as before stated, p. 91., with the superficial alluviums of Europe; and it was long before their real extent and thickness, and the various ages to which they belong, were fully recognized. They were observed to occur in patches, some of freshwater, others of marine origin, their geographical area being usually small as compared to the secondary formations, and their position often suggesting the idea of their having been deposited in different bays, lakes, estuaries, or inland seas, after a large portion of the space now occupied by Europe had already been converted into dry land. The first deposits of this class, of which the characters were accurately determined, were those occurring in the neighbourhood of Paris, described in 1810 by MM. Cuvier and Brongniart. They were ascertained to consist of successive sets of strata, some of marine, others of freshwater origin, lying one upon the other. The fossil shells and corals were perceived to be almost all of unknown Strata were soon afterwards brought to light in the vicinity of London, and in Hampshire, which, although dissimilar in mineral composition, were justly inferred by Mr. T. Webster to be of the same age as those of Paris, because the greater number of the fossil shells were specifically identical. For the same reason rocks found on the Gironde, in the South of France, and at certain points in the North of Italy, were suspected to be of contemporaneous origin. A variety of deposits were afterwards found in other parts of Europe, all reposing immediately on rocks as old or older than the chalk, and which exhibited certain general characters of resemblance in their organic remains to those previously observed near Paris and London. An attempt was therefore made at first to refer the whole to one period; and when at length this seemed impracticable, it was contended that as in the Parisian series there were many subordinate formations of considerable thickness which must have accumulated one after the other, during a great lapse of time, so the various patches of tertiary strata scattered over Europe might correspond in age, some of them to the older, and others to the newer, subdivisions of the Parisian series. This error, although most unavoidable on the part of those who made the first generalizations in this branch of geology, retarded seriously for some years the progress of classification. A more scrupulous attention to specific distinctions, aided by a careful regard to the relative position of the strata containing them, led at length to the conviction that there were formations both marine and freshwater of various ages, and all newer than the strata of the neighbourhood of Paris and London. One of the first steps in this chronological reform was made in 1811, by an English naturalist, Mr. Parkinson, who pointed out the fact that certain shelly strata, provincially termed "Crag" in Suffolk, lay decidedly over a deposit which was the continuation of the blue clay of London. At the same time he remarked that the fossil testacea in these newer beds were distinct from those of the blue clay, and that while some of them were of unknown species, others were identical with species now inhabiting the British seas. Another important discovery was soon afterwards made by Brocchi in Italy, who investigated the argillaceous and sandy deposits replete with shells which form a low range of hills, flanking the Apennines on both sides, from the plains of the Po to Calabria. These lower hills were called by him the Subapennines, and were formed of strata of different ages, all newer than those of Paris and London. Another tertiary group occurring in the neighbourhood of Bordeaux and Dax, in the south of France, was examined by M. de Basterot in Without pretending to give a complete sketch of the progress of discovery, I may refer to the facts above enumerated, as illustrating the course usually pursued by geologists when they attempt to found new chronological divisions. The method bears some analogy to that pursued by the naturalist in the construction of genera, when he selects a typical species, and then classes as congeners all other species of animals and plants which agree with this standard within certain limits. The genera A. and C. having been founded on these principles, a new species is afterwards met with, departing widely both from A. and C., but in many respects of an intermediate character. For this new type it becomes necessary to institute the new genus B., in which are included all species afterwards brought to light, which agree more nearly with B. than with the types of A. or C. In like manner a new formation is met with in geology, and the characters of its fossil fauna and flora investigated. From that moment it is considered as a record of a certain period of the earth's history, and a standard to which other deposits may be compared. If any are found containing the same or nearly the same organic remains, and occupying the same relative position, they are regarded in the light of contemporary annals. All such monuments are said to relate to one period, during which certain events occurred, such as the formation of particular rocks by aqueous or volcanic agency, or the continued existence and fossilization of certain tribes of animals and plants. When several of these periods have had their true places assigned to them in a chronological series, others are discovered which it becomes necessary to intercalate between those first known; and the difficulty of assigning clear lines of separation must unavoidably increase in proportion as chasms in the past history of the globe are filled up. Every zoologist and botanist is aware that it is a comparatively easy task to establish genera in departments which have been enriched The isolated position of distinct tertiary deposits in different parts of Europe has been already alluded to. In addition to the difficulty presented by this want of continuity when we endeavour to settle the chronological relations of these deposits, another arises from the frequent dissimilarity in mineral character of strata of contemporaneous date, such, for example, as those of London and Paris before mentioned. The identity or non-identity of species is also a criterion which often fails us. For this we might have been prepared, for we have already seen, that the Mediterranean and Red Sea, although within 70 miles of each other, on each side of the Isthmus of Suez, have each their peculiar fauna; and a marked difference is found in the four groups of testacea now living in the Baltic, English Channel, Black Sea, and Mediterranean, although all these seas have many species in common. In like manner a considerable diversity in the fossils of different tertiary formations, which have been thrown down in distinct seas, estuaries, bays, and lakes, does not always imply a distinctness in the times when they were produced, but may have arisen from climate and conditions of physical geography wholly independent of time. On the other hand, it is now abundantly clear, as the result of geological investigation, that different sets of tertiary strata, immediately superimposed upon each other, contain distinct imbedded species of fossils, in consequence of fluctuations which have been going on in the animate creation, and by which in the course of ages one state of things in the organic world has been substituted for another wholly dissimilar. It has also been shown that in proportion as the age of a tertiary deposit is more modern, so is its fauna more analogous to that now in being in the neighbouring seas. It is this law of a nearer agreement of the fossil testacea with the species now living, which may often furnish us with a clue for the chronological arrangement of scattered deposits, where we cannot avail ourselves of any one of the three ordinary chronological tests; namely, superposition, mineral character, and the specific identity of the fossils. To take another example, where the fossil fauna recedes a few steps farther back from our own times. We may compare, first, the beds of loam and clay bordering the Clyde in Scotland (called glacial by some geologists), secondly, others of fluvio-marine origin near Norwich, and, lastly, a third set often rising to considerable heights in Sicily, and we discover that in every case more than three-fourths of the shells agree with species still living, while the remainder are extinct. Hence we may conclude that all these, greatly diversified as are their organic remains, belong to one and the same era, or to a period immediately antecedent to the Post-Pliocene, because there has been time in each of the areas alluded to for an equal or nearly equal amount of change in the marine testaceous fauna. Contemporaneousness of origin is inferred in these cases, in spite of the most marked differences of mineral character or organic contents, from a similar degree of divergence in the shells from those now living in the adjoining seas. The advantage of such a test consists in supplying us with a common point of departure in all countries, however remote. But the farther we recede from the present times, and the smaller the relative number of recent as compared with extinct species in the tertiary deposits, the less confidence can we place in the exact value of such a test, especially when comparing the strata of very distant regions; for we cannot presume that the rate of former alterations in the animate world, or the continual going out and coming in of species, has been every where exactly equal in equal quantities of time. The form of the land and sea, and the climate, may have changed more in one region than in another; and consequently there may have been a more rapid destruction and renovation of species in one part of the globe than elsewhere. Considerations of this kind should undoubtedly put us on our guard against relying too implicitly on the accuracy of this test; yet it can never fail to throw great light on the chronological relations of tertiary groups with each other, and with the Post-Pliocene period. We may derive a conviction of this truth not only from a study of geological monuments of all ages, but also by reflecting on the tendency which prevails in the present state of nature to a uniform rate of simultaneous fluctuation in the flora and fauna of the whole globe. The grounds of such a doctrine cannot be discussed here, and I It will be observed that in the foregoing allusions to organic remains, the testacea or the shell-bearing mollusca are selected as the most useful and convenient class for the purposes of general classification. In the first place, they are more universally distributed through strata of every age than any other organic bodies. Those families of fossils which are of rare and casual occurrence are absolutely of no avail in establishing a chronological arrangement. If we have plants alone in one group of strata and the bones of mammalia in another, we can draw no conclusion respecting the affinity or discordance of the organic beings of the two epochs compared; and the same may be said if we have plants and vertebrated animals in one series and only shells in another. Although corals are more abundant, in a fossil state, than plants, reptiles, or fish, they are still rare when contrasted with shells, especially in the European tertiary formations. The utility of the testacea is, moreover, enhanced by the circumstance that some forms are proper to the sea, others to the land, and others to freshwater. Rivers scarcely ever fail to carry down into their deltas some land shells, together with species which are at once fluviatile and lacustrine. By this means we learn what terrestrial, freshwater, and marine species co-existed at particular eras of the past; and having thus identified strata formed in seas with others which originated contemporaneously in inland lakes, we Among other characters of the molluscous animals, which render them extremely valuable in settling chronological questions in geology, may be mentioned, first, the wide geographical range of many species; and, secondly, what is probably a consequence of the former, the great duration of species in this class, for they appear to have surpassed in longevity the greater number of the mammalia and fish. Had each species inhabited a very limited space, it could never, when imbedded in strata, have enabled the geologist to identify deposits at distant points; or had they each lasted but for a brief period, they could have thrown no light on the connection of rocks placed far from each other in the chronological, or, as it is often termed, vertical series. Many authors have divided the European tertiary strata into three groups—lower, middle, and upper; the lower comprising the oldest formations of Paris and London before-mentioned; the middle those of Bordeaux and Touraine; and the upper all those newer than the middle group. When engaged in 1828 in preparing my work on the Principles of Geology, I conceived the idea of classing the whole series of tertiary strata in four groups, and endeavouring to find characters for each, expressive of their different degrees of affinity to the living fauna. With this view, I obtained information respecting the specific identity of many tertiary and recent shells from several Italian naturalists, and among others from Professors Bonelli, Guidotti, and Costa. Having in 1829 become acquainted with M. Deshayes, of Paris, already well known by his conchological works, I learnt from him that he had arrived, by independent researches, and by the study of a large collection of fossil and recent shells, at very similar views respecting the arrangement of tertiary formations. At my request he drew up, in a tabular form, lists of all the shells known to him to occur both in some tertiary formation and in a living state, for the express purpose of ascertaining the proportional number of fossil species identical with the recent which characterized successive groups; and this table, planned by us in common, was published by me in 1833. The term Miocene (from e???, meion, less, and ?a????, cainos, recent) is intended to express a minor proportion of recent species (of testacea), the term Pliocene (from p?e???, pleion, more, and ?a????, cainos, recent) a comparative plurality of the same. It may assist the memory of students to remind them, that the Miocene contain a minor proportion, and Pliocene a comparative plurality of recent species; and that the greater number of recent species always implies the more modern origin of the strata. It has sometimes been objected to this nomenclature that certain species of infusoria found in the chalk are still existing, and, on the other hand, the Miocene and Older Pliocene deposits often contain the remains of mammalia, reptiles, and fish, exclusively of extinct species. But the reader must bear in mind that the terms Eocene, Miocene, and Pliocene were originally invented with reference purely to conchological data, and in that sense have always been and are still used by me. The distribution of the fossil species from which the results before mentioned were obtained in 1830 by M. Deshayes was as follows:—
Since the year 1830 the progress of conchological science has been most rapid, and the number of living species obtained from different parts of the globe has been raised from about 5000 to more than 10,000. New fossil species have also been added to our collections in great abundance; and at the same time a more copious supply of individuals both of fossil and recent species, some of which were previously very rare, have been procured, affording more ample data for determining the specific character. Besides the reforms introduced in consequence of these new zoological facilities, other errors of a geological nature have been in many instances removed. Post-Pliocene Formations.I have adopted the term Post-Pliocene for those strata which are sometimes called post-tertiary or modern, and which are characterized These modern formations, thus defined, comprehend not only those strata which can be shown to have originated since the earth was inhabited by man, but also deposits of far greater extent and thickness, in which no signs of man or his works can be detected. In some of these, of a date long anterior to the times of history and tradition, the bones of extinct quadrupeds have been met with of species which probably never co-existed with the human race, as, for example, the mammoth, mastodon, megatherium, and others, and yet the shells are the same as those now living. That portion of the post-pliocene group which belongs to the human epoch, and which is sometimes called Recent, forms a very unimportant feature in the geological structure of the earth's crust. I have shown, however, in "The Principles," where the recent changes of the earth illustrative of geology are described at length, that the deposits accumulated at the bottom of lakes and seas within the last 4000 or 5000 years can neither be insignificant in volume or extent. They lie hidden, for the most part, from our sight; but we have opportunities of examining them at certain points where newly-gained land in the deltas of rivers has been cut through during floods, or where coral reefs are growing rapidly, or where the bed of a sea or lake has been heaved up by subterranean movements and laid dry. Their age may be recognized either by our finding in them the bones of man in a fossil state, that is to say, imbedded in them by natural causes, or by their containing articles fabricated by the hands of man. Thus at Puzzuoli, near Naples, marine strata are seen containing fragments of sculpture, pottery, and the remains of buildings, together with innumerable shells retaining in part their colour, and of the same species as those now inhabiting the Bay of BaiÆ. The uppermost of these beds is about 20 feet above the level of the sea. Their emergence can be proved to have taken place since the beginning of the sixteenth century. In Ischia, a small island near Naples, composed in like manner of marine and volcanic formations, Dr. Philippi collected in the stratified tuff and clay ninety-two species of shells of existing species. In the centre of Ischia, the lofty hill called Epomeo, or San Nicola, is composed of greenish indurated tuff, of a prodigious thickness, interstratified in some parts with marl, and here and there with great beds of solid lava. Visconti ascertained by trigonometrical measurement that this mountain was 2605 feet above the level of the sea. Not far from its summit, at the height of about 2000 feet, as also near Moropano, a village only 100 feet lower, on the southern declivity of the mountain, I collected, in 1828, many shells of species now inhabiting the neighbouring gulf. It is clear, therefore, that the great mass of Epomeo was not only raised to its present height, but was also formed beneath the waters, within the post-pliocene period. It is a fact, however, of no small interest, that the fossil shells from these modern tuffs of the volcanic region surrounding the Bay of BaiÆ, although none of them extinct, indicate a slight want of correspondence between the ancient fauna and that now inhabiting the Mediterranean. Philippi informs us that when he and M. Scacchi had collected ninety-nine species of them, he found that only one, Pecten medius, now living in the Red Sea, was absent from the Mediterranean. Notwithstanding this, he adds, "the condition of the sea when the tufaceous beds were deposited must have been considerably different from its present state; for Tellina striata was then common, and is now rare; Lucina spinosa was both more abundant and grew to a larger size; Lucina fragilis, now rare, and hardly measuring 6 lines, then attained the enormous dimensions of 14 lines, and was extremely abundant; and Ostrea lamellosa, Broc., no longer met with near Naples, existed at that time, and attained a size so large that one lower valve has been known to measure 5 inches 9 lines in length, 4 inches in breadth, 11/2 inch in thickness, and weighed 261/2 ounces." There are other parts of Europe where no volcanic action manifests itself at the surface, as at Naples, whether by the eruption of lava or by earthquakes, and yet where the land and bed of the adjoining sea are undergoing upheaval. The motion is so gradual as to be insensible to the inhabitants, being only ascertainable by careful scientific measurements compared after long intervals. Such an upward movement has been proved to be in progress in Norway and Sweden throughout an area about 1000 miles N. and S., and for an unknown distance E. and W., the amount of elevation always increasing as we On the opposite or western coast of Sweden, at Uddevalla, post-pliocene strata, containing recent shells, not of that brackish water character peculiar to the Baltic, but such as now live in the northern ocean, ascend to the height of 200 feet; and beds of clay and sand of the same age attain elevations of 300 and even 700 feet in Norway, where they have been usually described as "raised beaches." They are, however, thick deposits of submarine origin, spreading far and wide, and filling valleys in the granite and gneiss, just as the tertiary formations, in different parts of Europe, cover or fill depressions in the older rocks. It is worthy of remark, that although the fossil fauna characterizing these upraised sands and clays consists exclusively of existing northern species of testacea, yet, according to LovÉn (an able living naturalist of Norway), the species do not constitute such an assemblage as now inhabits corresponding latitudes in the German Ocean. On the contrary, they decidedly represent a more arctic fauna. Judging by the uniformity of climate now prevailing from century to century, and the insensible rate of variation in the organic world in our own times, we may presume that an extremely lengthened period was required even for so slight a modification of the molluscous fauna, as that of which the evidence is here brought to light. On the other hand, we have every reason for inferring on independent grounds (namely, the rate of upheaval of land in modern times) that the antiquity of the deposits in question must be very great. For if In England, buried ships have been found in the ancient and now deserted channels of the Rother in Sussex, of the Mersey in Kent, and the Thames near London. Canoes and stone hatchets have been dug up, in almost all parts of the kingdom, from peat and shell-marl; but there is no evidence, as in Sweden, Italy, and many other parts of the world, of the bed of the sea, and the adjoining coast, having been uplifted bodily to considerable heights within the human period. Recent strata have been traced along the coasts of Peru and Chili, inclosing shells in abundance, all agreeing specifically with those now swarming in the Pacific. In one bed of this kind, in the island of San Lorenzo, near Lima, Mr. Darwin found, at the altitude of 85 feet above the sea, pieces of cotton-thread, plaited rush, and the head of a stalk of Indian corn, the whole of which had evidently been imbedded with the shells. At the same height on the neighbouring mainland, he found other signs corroborating the opinion that the ancient bed of the sea had there also been uplifted 85 feet, since the region was first peopled by the Peruvian race. In the West Indies, also, in the island of Guadaloupe, a solid limestone occurs, at the level of the sea-beach, enveloping human skeletons. The stone is extremely hard, and chiefly composed of comminuted shell and coral, with here and there some entire corals and shells, of species now living in the adjacent ocean. With them are included arrow-heads, fragments of pottery, and other articles of human workmanship. A limestone with similar contents has been formed, and is still forming, in St. Domingo. But there are also more ancient rocks in the West Indian Archipelago, as in Cuba, near the Havanna, and in other islands, in which are shells identical with those now living in corresponding latitudes; some well-preserved, others in the state of casts, all referable to the post-pliocene period. I have already described in the seventh chapter, p. 84., what would be the effects of oscillations and changes of level in any region drained by a great river and its tributaries, supposing the area to be first depressed several hundred feet, and then re-elevated. I believe that such changes in the relative level of land and sea have actually occurred in the post-pliocene era in the hydrographical basin of the Mississippi and in that of the Rhine. The accumulation of fluviatile matter in a delta during a slow subsidence may raise the newly Fig. 106. Valley of the Mississippi.
I have endeavoured to show that the deposits forming the delta and alluvial plain of the Mississippi consist of sedimentary matter, extending over an area of 30,000 square miles, and known in some parts to be several hundred feet deep. Although we cannot estimate correctly how many years it may have required for the river to bring down from the upper country so large a quantity of earthy matter—the data for such a computation being as yet incomplete—we may still approximate to a minimum of the time which such an operation must have taken, by ascertaining experimentally the annual discharge of water by the Mississippi, and the mean annual amount of solid matter contained in its waters. The lowest estimate of the time required would lead us to assign a high antiquity, amounting to many tens of Loess of the Valley of the Rhine.—A similar succession of geographical changes, attended by the production of a fluviatile formation, singularly resembling that which bounds the great plain of the Mississippi, seems to have occurred in the hydrographical basin of the Rhine, since the time when that basin had already acquired its present outline of hill and valley. I allude to the deposit provincially termed loess in part of Germany, or lehm in Alsace, filled with land and freshwater shells of existing species. It is a finely comminuted sand or pulverulent loam of a yellowish grey colour, consisting chiefly of argillaceous matter combined with a sixth part of carbonate of lime, and a sixth of quartzose and micaceous sand. It often contains calcareous sandy concretions or nodules, rarely exceeding the size of a man's head. Its entire thickness amounts, in some places, to between 200 and 300 feet; yet there are often no signs of stratification in the mass, except here and there at the bottom, where there is occasionally a slight intermixture of drifted materials derived from subjacent rocks. Unsolidified as it is, and of so perishable a nature, that every streamlet flowing over it cuts out for itself a deep gully, it usually terminates in a vertical cliff, from the surface of which land shells are seen here and there to project in relief. In all these features it presents a precise counterpart to the loess of the Mississippi. It is so homogeneous as generally to exhibit no signs of stratification, owing, probably, to its materials having been derived from a common source, and having been accumulated by a uniform action. Yet it displays in some few places decided marks of successive deposition, where coarser and finer materials alternate, especially near the bottom. Calcareous concretions, also enclosing land-shells, are sometimes arranged in horizontal layers. It is a remarkable deposit, from its position, wide extent, and thickness, its homogeneous mineral composition, and freshwater origin. Its distribution clearly shows that after the great valley of the Rhine, from Schaffhausen to Bonn, had acquired its present form, having its bottom strewed over with coarse gravel, a period arrived when it became filled up from side to side with fine mud, which was also thrown down in the valleys of the principal tributaries of the Rhine. Thus, for example, it may be traced far into WÜrtemberg, up the valley of the Neckar, and from Frankfort, up the valley of the Main, to above Dettelbach. I have also seen it spreading over the country of Mayence, Eppelsheim, and Worms, on the left bank of the Rhine, and on the opposite side on the table-land above the Bergstrasse, between But, in general, the loess overlies all the volcanic products, even those between Neuwied and Bonn, which have the most modern aspect; and it has filled up in part the crater of the Roderberg, an extinct volcano near Bonn. In 1833 a well was sunk at the bottom of this crater, through 70 feet of loess, in part of which were the usual calcareous concretions. The interstratification above alluded to, of loess with layers of pumice and volcanic ashes, has led to the opinion that both during and since its deposition some of the last volcanic eruptions of the Lower Eifel have taken place. Should such a conclusion be adopted, we should be called upon to assign a very modern date to these eruptions. This curious point, therefore, deserves to be reconsidered; since it may possibly have happened that the waters of the Rhine, swollen by the melting of snow and ice, and flowing at a great height through a valley choked up with loess, may have swept away the loose superficial scoriÆ and pumice of the Eifel volcanos, and spread them out occasionally over the yellow loam. Sometimes, also, the melting of snow on the slope of small volcanic cones may have given rise to local floods, capable of sweeping down light pumice into the adjacent low grounds. The first idea which has occurred to most geologists, after examining the loess between Mayence and Basle, is to imagine that a great lake once extended throughout the valley of the Rhine between those two places. Such a lake may have sent off large branches up the course of the Main, Neckar, and other tributary valleys, in all of which large patches of loess are now seen. The barrier of the lake might be placed somewhere in the narrow and picturesque gorge of the Rhine between Bingen and Bonn. But this theory fails altogether to explain the phenomena; when we discover that that gorge itself has once been filled with loess, which must have been tranquilly deposited in it, as also in the lateral valley of the Lahn, communicating with the gorge. The loess has also overspread the high adjoining platform near the village of Plaidt above Andernach. Nay, on proceeding farther down to the north, we discover that the Besides these objections to the lake theory, the loess is met with near Basle, capping hills more than 1200 feet above the sea; so that a barrier of land capable of separating the supposed lake from the ocean would require to be, at least, as high as the mountains called the Siebengebirge, near Bonn, the loftiest summit of which, the Oehlberg, is 1209 feet above the Rhine and 1369 feet above the sea. It would be necessary, moreover, to place this lofty barrier somewhere below Cologne, or precisely where the level of the land is now lowest. Instead, therefore, of supposing one continuous lake of sufficient extent and depth to allow of the simultaneous accumulation of the loess, at various heights, throughout the whole area where it now occurs, I formerly suggested that, subsequently to the period when the countries now drained by the Rhine and its tributaries had nearly acquired their actual form and geographical features, they were again depressed gradually by a movement like that now in progress on the west coast of Greenland. The proportion of land shells of the genera Helix, Pupa, and Bulimus, is very large in the loess; but in many places aquatic species of the genera Lymnea, Paludina, and Planorbis are also found. These may have been carried away during floods from shallow pools and marshes bordering the river; and the great extent of marshy ground caused by the wide overflowings of rivers above supposed would favour the multiplication of amphibious mollusks, such as the Succinea (fig. 107.), which is almost everywhere characteristic of this formation, and is sometimes accompanied, as near Bonn, by another species, S. amphibia (fig. 34. p. 29.). Among other abundant Fig. 107. Succinea elongata. Fig. 108. Pupa muscorum. Fig. 109. Helix plebeium. Bones of vertebrated animals are rare in the loess, but those of the mammoth, horse, and some other quadrupeds have been met with. At the village of Binningen, and the hills called Bruderholz, near Basle, I found the vertebrÆ of fish, together with the usual shells. These vertebrÆ, according to M. Agassiz, belong decidedly to the Shark family, perhaps to the genus Lamna. In explanation of their occurrence among land and freshwater shells, it may be stated that certain fish of this family ascend the Senegal, Amazon, and other great rivers, to the distance of several hundred miles from the ocean. At Cannstadt, near Stuttgart, in a valley also belonging to the hydrographical basin of the Rhine, I have seen the loess pass downwards into beds of calcareous tuff and travertin. Several valleys in northern Germany, as that of the Ilm at Weimar, and that of the Tonna, north of Gotha, exhibit similar masses of modern limestone filled with recent shells of the genera Planorbis, Lymnea, Paludina, &c., from 50 to 80 feet thick, with a bed of loess much resembling that of the Rhine, occasionally incumbent on them. In these modern limestones used for building, the bones of Elephas primigenius, Rhinoceros tichorinus, Ursus spelÆus, HyÆna spelÆa, with the horse, ox, deer, and other quadrupeds, occur; and in 1850 Mr. H. Credner and I obtained in a quarry at Tonna, at the depth of 15 feet, inclosed in the calcareous rock and surrounded with dicotyledonous leaves and petrified leaves, four eggs of a snake of the size of the largest European Coluber, which, with three others, had been found lying in a series, or string. They are, I believe, the first reptilian remains which have been met with in strata of this age. The agreement of the shells in these cases with recent European species enables us to refer to a very modern period the filling up and re-excavation of the valleys; an operation which doubtless consumed a long period of time, since which the mammiferous fauna has undergone a considerable change. |