If we make a thin slice across the stem of a rapidly growing plant,—e.g. geranium, begonia, celery,—mount it in water, and examine it microscopically, it will be found to be made up of numerous cavities or chambers separated by delicate partitions. Often these cavities are of sufficient size to be visible to the naked eye, and examined with a hand lens the section appears like a piece of fine lace, each mesh being one of the chambers visible when more strongly magnified. These chambers are known as “cells,” and of them the whole plant is built up.

In order to study the structure of the cell more exactly we will select such as may be examined without cutting them. A good example is furnished by the common spiderwort (Fig.1). Attached to the base of the stamens (Fig.85, B) are delicate hairs composed of chains of cells, which may be examined alive by carefully removing a stamen and placing it in a drop of water under a cover glass. Each cell (Fig.1) is an oblong sac, with a delicate colorless wall which chemical tests show to be composed of cellulose, a substance closely resembling starch. Within this sac, and forming a lining to it, is a thin layer of colorless matter containing many fine granules. Bands and threads of the same substance traverse the cavity of the cell, which is filled with a deep purple homogeneous fluid. This fluid, which in most cells is colorless, is called the cell sap, and is composed mainly of water. Imbedded in the granular lining of the sac is a roundish body (n), which itself has a definite membrane, and usually shows one or more roundish bodies within, besides an indistinctly granular appearance. This body is called the nucleus of the cell, and the small one within it, the nucleolus.

The membrane surrounding the cell is known as the cell wall, and in young plant cells is always composed of cellulose.

The granular substance lining the cell wall (Fig.1, pr.) is called “protoplasm,” and with the nucleus constitutes the living part of the cell. If sufficiently magnified, the granules within the protoplasm will be seen to be in active streaming motion. This movement, which is very evident here, is not often so conspicuous, but still may often be detected without difficulty.

The cell may be regarded as the unit of organic structure, and of cells are built up all of the complicated structures of which the bodies of the highest plants and animals are composed. We shall find that the cells may become very much modified for various purposes, but at first they are almost identical in structure, and essentially the same as the one we have just considered.

Very many of the lower forms of life consist of but a single cell which may occasionally be destitute of a cell wall. Such a form is shown in Figure2. Here we have a mass of protoplasm with a nucleus (n) and cavities (vacuoles, v) filled with cell sap, but no cell wall. The protoplasm is in constant movement, and by extensions of a portion of the mass and contraction of other parts, the whole creeps slowly along. Other naked cells (Fig.12, B; Fig.16, C) are provided with delicate thread-like processes of protoplasm called “cilia” (sing. cilium), which are in active vibration, and propel the cell through the water.

On placing a cell into a fluid denser than the cell sap (e.g. a ten-per-cent solution of sugar in water), a portion of the water will be extracted from the cell, and we shall then see the protoplasm receding from the wall (Fig.4, C), showing that it is normally in a state of tension due to pressure from within of the cell sap. The cell wall shows the same thing though in a less degree, owing to its being much more rigid than the protoplasmic lining. It is owing to the partial collapsing of the cells, consequent on loss of water, that plants wither when the supply of water is cut off.

As cells grow, new ones are formed in various ways. If the new cells remain together, cell aggregates, called tissues, are produced, and of these tissues are built up the various organs of the higher plants. The simplest tissues are rows of cells, such as form the hairs covering the surface of the organs of many flowering plants (Fig.3), and are due to a division of the cells in a single direction. If the divisions take place in three planes, masses of cells, such as make up the stems, etc., of the higher plants, result (Fig.4, A, B).