The manner in which these forests were fossilized may next be considered. Though the whole history of the process is not fully understood, it was undoubtedly dependent on or at least greatly facilitated by the presence of volcanic and hydrothermal activity, The fossil forests are surrounded by a matrix that is known as an acidic lava—that is, a siliceous lava—which contains abundant silica in solution. The first part of the process of silicification may well have been that above described as taking place in the hot spring areas at the present day—that is, the silica would be deposited in all the cells and vessels of the wood, making an accurate cast of all open spaces. Then, while the slow process of decay went on, as each particle of organic matter was removed its place was taken by the silica, until, finally, all the wood substance had disappeared and its place atom by atom had been taken by silica. By this or a similar process the wood has been preserved or fossilized with remarkable fidelity: in fact, thin sections or slices of the fossil wood may be studied under the higher powers of the microscope with almost or quite as much completeness and satisfaction as if they were sections cut from a piece of living wood. Each cell and vessel, with its characteristic pits and markings, is preserved exactly as it grew. Some of the wood, however, was evidently more or less decayed before it was fossilized, or else decay worked faster than replacement, so that in some fragments the structure is not so clearly preserved. Many of the trunks were subjected to pressure before replacement was complete, and this has crushed or distorted the cells. On the whole, however, the wood is exceptionally well preserved, as may be seen in figures 12, 13, 14, and 15. These are all magnified 100 diameters and were photographed directly from the thin sections—that is, they are photomicrographs—and have not been retouched in any manner. Figure 12 shows a transverse section of the wood of the large redwood trunk that has been so often mentioned (see title page). The section is cut through one of the growth rings, which consists of 12 or 15 rows of very thick-walled cells. The large, regular thin-walled cells, which begin abruptly above the growth ring, belong to the spring wood—that is, the wood first formed after growth starts in spring, when the supply of nourishment is abundant. If there is sufficient moisture and all conditions are favorable this vigorous growth of wood cells may continue without interruption until the approach of cold or dry weather, but not infrequently there may be a brief shortage of moisture, and this is reflected in the formation of a few rows of thicker-walled cells. Such a condition may be observed in the present specimen, in which a slight, partial ring may be seen at some distance above the main ring. Fig. 12.—Thin section of wood of fossil redwood (Sequoia magnifica), showing growth ring. Section transverse. Magnified 100 diameters. Fig. 13.—Thin section of wood of fossil pine (Pityoxylon amethystinum), showing growth ring and resin tube. Section transverse. Magnified 100 diameters. Fig. 14.—Thin section of wood of fossil pine (Pityoxylon aldersoni), showing medullary rays and resin tube. Section tangential. Magnified 100 diameters. Fig. 15.—Thin section of wood of fossil laurel (Laurinoxylon pulchrum), showing wood cells, tubes, and rays. Section longitudinal. Magnified 100 diameters. The very perfect preservation of the wood of one of the pines (Pityoxylon amethystinum) is shown in figure 13, a view of a section cut through a part of a growth ring and into the spring and summer wood, the rings in this species being so broad that it is impossible to show a complete one. The opening near the bottom of the figure shows one of the large resin ducts, which, in the living wood, is filled with the “pitch” that so readily exudes when a branch is cut or broken. A longitudinal section of the other species of pine (Pityoxylon aldersoni) is shown in figure 14. The many little rows of superimposed cells in the midst of the long wood cells are the cut-off ends of what are known as medullary rays—that is, the little plates of cells that connect pith and bark. One of the resin cells cut in the long direction is shown near the center of the figure; the contents are much darker than that of the wood cells. The very great difference between the sections of coniferous wood just described and the wood of a deciduous tree is brought out in figure 15, which is a longitudinal section of a laurel (Laurinoxylon pulchrum). In this the wood cells are relatively much smaller and shorter, and the medullary rays are in several irregular rows. The large dotted duct near the middle of the figure is a feature not present in coniferous trees. |