CHAPTER XI. HABITS AND HAIR OF CARNIVORES. Cats.

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Another large and important order of hair-clad mammals must now be considered, and the same course as in the case of the ungulates will be followed; the two leading families of FelidÆ and CanidÆ will be taken, and a type of each examined in reference to its hair-distribu­tion. Lydekker gives about 100 pages to the cats and 80 to the dogs, so from the point of view of general biology there seems little to choose between them. The bears, racoons, weasel tribe, seals and walruses may be put out of account. They are painfully old-fashioned or Normal as to the arrangement of their hair.

First things first is always a good rule, and there is little doubt where we ought to begin among the families and species Carnivores. Among FelidÆ one cannot unfortunately choose the harmless necessary cat of tiles, areas, firesides and ladies’ laps, to say nothing of those lovers of cats like Huxley who would never eject his cat from his armchair if she had been there before him. It is true that we know much of her daily and nightly mode of life—many of us too much—and in that respect one could set to work with confidence in interpreting her hair patterns, but on account of her long and thick coat we can only speculate what patterns or innovations of her family uniform she might have devised; but here we are not concerned with romance or the “might have beens.” It will be remarked that one perforce unconsciously calls the domestic cat “she” as sailors do their ships. I understand that in Somersetshire they call everything of their common life “he” except the tom-cat who is always “she.” The reasons for the use of genders in different creatures would be an interesting little study.

Lion.

The King of Beasts will, therefore, be the hero of this chapter. Lydekker tells us that the lion, like many heroes of antiquity who are no heroes to their valets, in spite of his character for grandeur, nobility and courage, has been subjected to the merciless higher criticism of modern travellers, Selous, Livingstone, and others, and he has been shown up as cowardly by nature and mean in his general conduct. It remains for some learned scholar to whitewash the hyÆna, as someone has done for Caesar Borgia, and to put him in the place of the lion. But Lydekker does not admit that this disparagement of the lion goes very far. He is the King of Beasts by grandeur of appearance, strength and ferocity.

Fig. 36.—Lioness, showing by arrows the direction of hair-streams on muzzle, parting from one another at the level of the orbits.

The lion’s skin is covered by close fine hair, except in certain seasons in cold climates, and is easily studied. There are three regions where this representative cat has departed from the Primitive mammalian slope of hair, and the figure of a lioness shows two of these, the peculiar downward trend of hair on the muzzle and the whorl on the shoulder. Fig.37 shows the third, A C, on the middle of the back as well as the whorls at D.

Snout of the Cats

The muzzle of all the cats is very short and broad, and at the level of the orbits shows a peculiar reversal of the hair from the rest of the head, for instead of being like that of a dog in which the hair slopes all the way upwards from the tip of the snout to the rest of the head, it breaks away from this normal type and passes in a uniform close stream to the edge of the wet muzzle. The arrows in Fig.36 show this change. One asks at once the reason for such an unexpected trend of the hair on a small area, when the carnivores in other groups have a uniform slope towards the head from their more pointed muzzles. The cats have discarded the earlier family pattern and for a reason which does credit to their self-respect. Very few naturalists know, or have described so well the meticulous care which animals take of their coats, as Miss Frances Pitt did in the National Review, where she gave a delightful account of “How Animals Clean Themselves.” The toilet of the lion she did not discuss, perhaps for prudential reasons. Her account dealt chiefly with a number of small hairy mammals and lower forms of life. Watch a dog cleaning his coat and you will see the ingenious way in which he pushes his head and body forward as he lies on some rough surface such as grass, or our best drawing-room mat. He can thus clean his snout and other parts, but no cat adopts so rough and ready a method. We know how long and how scrupulously she licks her fur to clean it in the parts she can reach and cleans her head with her paws. But with such a broad snout as she and the larger cats possess she cannot clean the short surface of it in the manner of the dog. So she “dresses” this little surface in a special way of rubbing it from the neighbourhood of her eyes forward with her paws. And so we may assume does the chieftain of her clan finish off this little bit of his toilet. We are so much accustomed to dwell on the naturally clean habits of a domestic cat that without such an account as Miss Frances Pitt has given we should have hesitated to transfer the character for personal cleanliness from the domesticated to the wild cat. If this be not the sole reason for the course of the hair-stream I have described, I am at a loss to imagine any other.

Lion’s Neck.

On each side of the lion’s neck where it joins the shoulder there is a well-developed whorl, and this as a rule is extended forwards into a feathering (Figs.36 and 37), and ends in a crest on the lower part of the side of the neck. It is common also in tigers and leopards. This is, as elsewhere, a record of strong and oft-repeated action in powerful muscles which lie beneath it, and bears witness to the great functional activity of the fore-limbs as compared with the hind-limbs in these three formidable cats. It is not an animal pedometer, but may perhaps be termed an ergograph.

Lion’s Back.

Fig. 37.—Back of Lion, showing reversed area of hair with whorl at A. Feather­ing B. Crest C.

The strange pattern of reversed hair (Fig.37) is much the most notable of the three peculiarities found on the lion’s skin. It consists of a whorl (A) lying over the lumbar region in the middle line which expands into a very broad feathering (B) and terminates in a crest (C) a short distance behind the level of the shoulders. This is not found in any of the numerous short-haired FelidÆ that I have examined, and it is a feature which demands explana­tion. I know no other mammal, ungulate or carnivore, that has any pattern resembling this; indeed, if one were to photograph the pattern in question and a few inches of the skin surrounding it, and be told that it came from the back of a mammal one could not doubt that it was a hall-mark of the King of Beasts. It would not produce that thrill of intense interest which we felt at the meeting on 7th May, 1901, at the Zoological Society of London, when from a water colour sketch and three pieces of skin taken from the body of a hitherto unknown mammal, Sir Harry Johnston proceeded to reconstruct the Okapi, at first dubbed knight, as a member of the EquidÆ, but later promoted downwards to the GiraffidÆ. But one could do no less, with some knowledge of the hair of mammals, than reconstruct from such a photograph a large, powerful and ferocious carnivore, and where but in the lion can the greatest example of those attributes be found? I say this advisedly, for this remarkable pattern of the lion’s back is as much a stamp of his moral or mental quality as the Inguinal Pedometer is of the locomotive rÔle in life of equus caballus.

I hear the sharp voice of the critic here, “Come, come, you may have shown reason for the latter, but how on earth do mental and moral qualities of an animal come into your scheme?” Well, we have in this pattern of the lion’s back to deal with a unique phenomenon for the produc­tion of which neither pressure, nor friction, nor gravita­tion, nor underlying muscular traction will account. Nevertheless, it is a result of muscular action of a rare kind. Who does not know the striking appearance of the hair along the centre of a short-haired dog when he bristles up with rage or fear, or both combined, at the sight of a foe? This common event has its own mechanical cause, though it is one strictly governed by the mental and moral qualities of the dog, and we see the vivid proof before us of the action of the minute arrectores pili, in this particular region of the dog. It is precisely in the same situation that the special pattern of the lion’s hair is found. It is not for nothing that Nature has provided every tiny hair of the mammalian skin with that insignificant little band of muscle which lies within the hair-pit, and is attached to the sloping hair on its posterior side, and thus when it contracts serves to drag it into an erect position. I refrain from discussing what may be held to be the survival value, under the theory of selection, of this power of the arrectores pili to confer on the possessor an added appearance of ferocity and general frightfulness. This is quite a likely explana­tion of the presence of these little muscles. Be that as it may the modus operandi of the reversed hair which has become fixed on the lion’s back is made clear, theory of origin apart. And I submit that the presence of it in this region in this animal is a stamp of his persistently ferocious nature, as much as the various peculiarities of arrangement of hair on man’s eyebrows in a previous chapter are of the mental and moral habits of the individual man. As rulers of old used, in their genial fashion, to brand a supposed or actual criminal on his shoulder or forehead, so is the lion branded with an hereditary mark of his nature and the past life of himself and his ancestors. I doubt not that if short-haired terriers were living a wild life among numerous foes their bristling hair would have become fixed in a similar fashion. I would only here draw the attention of the reader to the fact that this reversed area of hair on the lion’s back cannot be held to add to the general frightfulness of the possessor. It would be invisible to an approaching foe, as it lies hidden behind the great head and mane. This pattern on the lion’s back will be referred to later in a somewhat different connec­tion.


                                                                                                                                                                                                                                                                                                           

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