A. H. Keane

The World peopled by Migration from one Centre by Pleistocene Man—The Primary Groups evolved each in its special Habitat—Pleistocene Man: Pithecanthropus erectus; The Mauer jaw, Homo Heidelbergensis; The Piltdown skull, Eoanthropus Dawsoni—General View of Pleistocene Man—The first Migrations—Early Man and his Works—Classification of Human Types: H. primigenius, Neandertal or Mousterian Man; H. recens, Galley Hill or Aurignacian Man—Physical Types—Human Culture: Reutelian, Mafflian, Mesvinian, Strepyan, Chellean, Acheulean, Mousterian, Aurignacian, Solutrian, Magdalenian, Azilian—Chronology—The early History of Man a Geological Problem—The Human Varieties the Outcome of their several Environments—Correspondence of Geographical with Racial and Cultural Zones.

In order to a clear understanding of the many difficult questions connected with the natural history of the human family, two cardinal points have to be steadily borne in mind—the specific unity of all existing varieties, and the dispersal of their generalised precursors over the whole world in pleistocene times. As both points have elsewhere been dealt with by me somewhat fully[1], it will here suffice to show their direct bearing on the general evolution of the human species from that remote epoch to the present day.

It must be obvious that, if man is specifically one, though not necessarily sprung of a single pair, he must have had, in homely language, a single cradle-land, from which the peopling of the earth was brought about by migration, not by independent developments from different species in so many independent geographical areas.

It follows further, and this point is all-important, that, since the world was peopled by pleistocene man, it was peopled by a generalised proto-human form, prior to all later racial differences. The existing groups, according to this hypothesis, have developed in different areas independently and divergently by continuous adaptation to their several environments. If they still constitute mere varieties, and not distinct species, the reason is because all come of like pleistocene ancestry, while the divergences have been confined to relatively narrow limits, that is, not wide enough to be regarded zoologically as specific differences.

The battle between monogenists and polygenists cannot be decided until more facts are at our disposal, and much will doubtless be said on both sides for some time to come[2]. Among the views of human origins brought forward in recent years should be mentioned the daring theory of Klaatsch[3]. Recognising two distinct human types, Neandertal and Aurignac (see pp. 8, 9 below), and two distinct anthropoid types, gorilla and orang-utan, he derives Neandertal man and African gorilla from one common ancestor, and Aurignac man and Asiatic orang-utan from another. Though anatomists, especially those conversant with anthropoid structure[4], are not able to accept this view, they admit that many difficulties may be solved by the recognition of more than one primordial stock of human ancestors[5]. The questions of adaptation to climate and environment[6], the possibilities of degeneracy, the varying degrees of physiological activity, of successful mutations, the effects of crossing and all the complicated problems of heredity are involved in the discussion, and it must be acknowledged that our information concerning all of these is entirely inadequate.

Nevertheless all speculations on the subject are not based merely on hypotheses, and three discoveries of late years have provided solid facts for the working out of the problem.

These discoveries were the remains of Pithecanthropus erectus[7] in Java, in 1892, of the Mauer jaw[8], near Heidelberg, in 1907, and of the Piltdown skull[9] in Sussex in 1912. Although the Mauer jaw was accepted without hesitation, the controversy concerning the correct interpretation of the Javan fossils has been raging for more than twenty years and shows no sign of abating, while Eoanthropus Dawsoni is too recent an intruder into the arena to be fairly dealt with at present. Certain facts however stand out clearly. In late pliocene or early pleistocene times certain early ancestral forms were already in existence which can scarcely be excluded from the Hominidae. In range they were as widely distributed as Java in the east to Heidelberg and Sussex in the west, and in spite of divergence in type a certain correlation is not impossible, even if the Piltdown specimen should finally be regarded as representing a distinct genus[10]. Each contributes facts of the utmost importance for the tracing out of the history of human evolution. Pithecanthropus raises the vexed question as to whether the erect attitude or brain development came first in the story. The conjunction of pre-human braincase with human thighbone appeared to favour the popular view that the erect attitude was the earlier, but the evidence of embryology suggests a reverse order. And although at first the thighbone was recognised as distinctly human it seems that of late doubts have been cast on this interpretation[11], and even the claim to the title erectus is called in question. The characters of straightness and slenderness on which much stress was laid are found in exaggerated form in gibbons and lemurs. The intermediate position in respect of mental endowment (in so far as brain can be estimated by cranial capacity) is shown in the accompanying diagram in which the cranial measurements of Pithecanthropus are compared with those of a chimpanzee and prehistoric man. The teeth strengthen the evidence, for they are described as too large for a man and too small for an ape. Thus Pithecanthropus has been confidently assigned to a place in a branch of the human family tree.

The Mauer jaw, the geological age of which is undisputed, also represents intermediate characters. The extraordinary strength and thickness of bone, the wide ascending ramus with shallow sigmoid notch (distinctly simian features) and the total absence of chin[12] would deny it a place among human jaws, but the teeth, which are all fortunately preserved in their sockets, are not only definitely human, but show in certain peculiarities less simian features than are to be found in the dentition of modern man[13].

The cranial capacity of the Piltdown skull, though variously estimated[14], is certainly greater than that of Pithecanthropus, the general outlines with steeply rounded forehead resemble that of modern man, and the bones are almost without exception typically human. The jaw, however, though usually attributed to the same individual[15], recalls the primitive features of the Mauer specimen in its thick ascending portion and shallow notch, while in certain characters it differs from any known jaw, ancient or modern[16]. The evidence afforded by the teeth is even more striking. The teeth of Pithecanthropus and of Homo Heidelbergensis were recognised as remarkably human, and although primitive in type, are far more advanced in the line of human evolution than the lowly features with which they are associated would lead one to expect. The Piltdown teeth are more primitive in certain characters than those of either the Javan or the Heidelberg remains. The first molar has been compared to that of Taubach, the most ape-like of human or pre-human teeth hitherto recorded, but the canine tooth (found by P. Teilhard in the same stratum in 1913[17]) finds no parallel in any known human jaw; it resembles the milk canine of the chimpanzee more than that of the adult dentition.

It cannot be said that any clear view of pleistocene man can be obtained from these imperfect scraps of evidence, valuable though they are. Rather may we agree with Keith that the problem grows more instead of less complex. "In our first youthful burst of Darwinianism we pictured our evolution as a simple procession of forms leading from ape to man. Each age, as it passed, transformed the men of the time one stage nearer to us—one more distant from the ape. The true picture is very different. We have to conceive an ancient world in which the family of mankind was broken up into narrow groups or genera, each genus again divided into a number of species—much as we see in the monkey or ape world of to-day. Then out of that great welter of forms one species became the dominant form, and ultimately the sole surviving one—the species represented by the modern races of mankind[18]."

We may assume therefore that the earth was mainly peopled by the generalised pleistocene precursors, who moved about, like the other migrating faunas, unconsciously, everywhere following the lines of least resistance, advancing or receding, and acting generally on blind impulse rather than of any set purpose.

That such must have been the nature of the first migratory movements will appear evident when we consider that they were carried on by rude hordes, all very much alike, and differing not greatly from other zoological groups, and further that these migrations took place prior to the development of all cultural appliances beyond the ability to wield a broken branch or a sapling, or else chip or flake primitive stone implements[19].

Herein lies the explanation of the curious phenomenon, which was a stumbling-block to premature systematists, that all the works of early man everywhere present the most startling resemblances, affording absolutely no elements for classification, for instance, during the times corresponding with the Chellean or first period of the Old Stone Age. The implements of palaeolithic type so common in parts of South India, South Africa, the Sudan, Egypt, etc., present a remarkable resemblance to one another. This, while affording a prima facies case for, is not conclusive of, the migrations of a definite type of humanity.

After referring to the identity of certain objects from the Hastings kitchen-middens and a barrow near Sevenoaks, W. J. L. Abbot proceeds: "The first thing that would strike one in looking over a few trays of these implements is the remarkable likeness which they bear to those of Dordogne. Indeed many of the figures in the magnificent 'Reliquiae Aquitanicae' might almost have been produced from these specimens[20]." And Sir J. Evans, extending his glance over a wider horizon, discovers implements in other distant lands "so identical in form and character with British specimens that they might have been manufactured by the same hands.... On the banks of the Nile, many hundreds of feet above its present level, implements of the European types have been discovered, while in Somaliland, in an ancient river valley, at a great elevation above the sea, Seton-Karr has collected a large number of implements formed of flint and quartzite, which, judging from their form and character, might have been dug out of the drift-deposits of the Somme and the Seine, the Thames or the ancient Solent[21]."

It was formerly held that man himself showed a similar uniformity, and all palaeolithic skulls were referred to one long-headed type, called, from the most famous example, the Neandertal, which was regarded as having close affinities with the present Australians. But this resemblance is shown by Boule[22] and others to be purely superficial, and recent archaeological finds indicate that more than one racial type was in existence in the Palaeolithic Age.

W. L. H. Duckworth on anatomical evidence constructs the following table[23].

H. Obermaier[24] argues as follows: Homo primigenius is neither the representative of an intermediate species between ape and man, nor a lower or distinct type than Homo sapiens, but an older primitive variety (race) of the latter, which survives in exceptional cases down to the present day[25]. Clearly then, according to the rules of zoological classification, we must term the two, Homo sapiens var. primigenius, as compared with Homo sapiens var. recens.

Whatever classification or nomenclature may be adopted the dual division in palaeolithic times is now generally recognised. The more primitive type is commonly called Neandertal man, from the famous cranium found in the Neandertal cave in 1857, or Mousterian man, from the culture associations. To this group belong the Gibraltar skull[26], and the skeletons from Spy[27], and Krapina, Croatia[28], together with the later discoveries (1908-11) at La Chapelle[29] (CorrÈze), Le Moustier[30], La Ferassie[31] (Dordogne) and many others.

Palaeolithic examples of the modern human type have been found at BrÜx (Bohemia)[32], BrÜnn (Moravia)[33] and Galley Hill in Kent[34], but the most complete find was that at Combe Capelle in 1909[35]. The numerous skeletons found at Cro-Magnon[36] and at the Grottes de Grimaldi at Mentone[37] though showing certain skeletal differences may be included in this group, the earliest examples of which are associated with Aurignacian culture[38].

From the evidence contributed by these examples the main characteristics of the two groups may be indicated, although, owing to the imperfection of the records, any generalisations must necessarily be tentative and subject to criticism.

The La Chapelle skull recalls many of the primitive features of the "ancestral types." The low receding forehead, the overhanging brow-ridges, forming continuous horizontal bars of bone overshadowing the orbits, the inflated circumnasal region, the enormous jaws, with massive ascending ramus, shallow sigmoid notch, "negative" chin and other "simian" characters seem reminiscent of Pithecanthropus and Homo Heidelbergensis. The cranial capacity however is estimated at over 1600 c.c., thus exceeding that of the average modern European, and this development, even though associated, as M. Boule has pointed out, with a comparatively lowly brain, is of striking significance. The low stature, probably about 1600 mm. (under 5½ feet) makes the size of the skull and cranial capacity all the more remarkable. "A survey of the characters of Neanderthal man—as manifested by his skeleton, brain cast, and teeth—have convinced anthropologists of two things: first, that we are dealing with a form of man totally different from any form now living; and secondly, that the kind of difference far exceeds that which separates the most divergent of modern human races[39]."

The earliest complete and authentic example of "Aurignacian man" was the skeleton discovered near Combe Capelle (Dordogne) in 1909[40]. The stature is low, not exceeding that of the Neandertal type, but the limb bones are slighter and the build is altogether lighter and more slender. The greatest contrast lies in the skull. The forehead is vertical instead of receding, and the strongly projecting brow-ridges are diminished, the jaw is less massive and less simian with regard to all the features mentioned above. Especially is this difference noticeable in the projection of the chin, which now for the first time shows the modern human outline. In short there are no salient features which cannot be matched among the living races of the present day.

On the cultural side no less than on the physical, the thousands of years which the lowest estimate attributes to the Early Stone Age were marked by slow but continuous changes.

The Reutelian (at the junction of the Pliocene and Pleistocene), Mafflian and Mesvinian industries, recognised by M. Rutot in Belgium, belong to the doubtful Eolithic Period, not yet generally accepted[41].

The lowest palaeolithic deposit is the Strepyan, so called from StrÉpy, near Charleroi, typically represented at St Acheul, Amiens, and recognised also in the Thames Valley[42]. The tools exhibit deliberate flaking, and mark the transition between eolithic and palaeolithic work. The associated fauna includes two species of elephant, E. meridionalis and E. antiquus, two species of rhinoceros, R. Etruscus and R. Merckii, and the hippopotamus. It is possible that the Mauer jaw and the Piltdown skull belong to this stage.

The Chellean industry[43], with the typical coarsely flaked almond-shaped implements, occurs abundantly in the South of England and in France, less commonly in Belgium, Germany, Austria-Hungary and Russia, while examples have been recognised in Palestine, Egypt, Somaliland, Cape Colony, Madras and other localities, though outside Europe the date is not always ascertainable and the form is not an absolute criterion[44].

Acheulean types succeed apparently in direct descent but the implements are altogether lighter, sharper, more efficient, and are characterised by finer workmanship and carefully retouched edges. A small finely finished lanceolate implement is typical of the sub-industry or local development at La Micoque (Dordogne).

The Chellean industry is associated with a warm climate and the remains of Elephas antiquus, Rhinoceros Merckii and hippopotamus. Lower Acheulean shows little variation, but with Upper Acheulean certain animals indicating a colder climate make their appearance, including the mammoth, Elephas primigenius, and the woolly rhinoceros, R. tichorhinus, but no reindeer.

The Mousterian industry is entirely distinct from its predecessors. The warm fauna has disappeared, the reindeer first occurs together with the musk ox, arctic fox, the marmot and other cold-loving animals. Man appears to have sought refuge in the caves, and from complete skeletons found in cave deposits of this stage we gain the first clear ideas concerning the physical type of man of the early palaeolithic period. Typical Mousterian implements consist of leaf-like or triangular points made from flakes struck from the nodule instead of from the dressed nodule itself, as in the earlier stages. The Levallois flakes, occurring at the base of the Mousterian (sometimes included in the Acheulean stage), initiate this new style of workmanship, but the Mousterian point shows an improvement in shape and a greater mastery in technique, producing a more efficient tool for piercing and cutting. Scrapers, carefully retouched, with a curved edge are also characteristic, besides many other forms. The complete skeletons from Le Moustier itself, La Chapelle, La Ferassie, and Krapina all belong to this stage, which marks the end of the lower palaeolithic period, the Age of the Mammoth.

The upper palaeolithic or Reindeer Age is divided into Aurignacian, Solutrian, and Magdalenian[45] culture stages, with the Azilian[46] separating the Magdalenian from the neolithic period. Each stage is distinguished by its implements and its art. The Aurignacian fauna, though closely resembling the Mousterian, indicates an amelioration of climate, the most abundant animals being the bison, horse, cave lion, and cave hyena, and human settlements are again found in the open. Among the typical implements are finely worked knife-like blades (ChÂtelperron point, Gravette point), keeled scrapers (TartÉ type), burins or gravers, and various tools and ornaments of bone. Art is represented by engravings and wall paintings, and to this stage belong statuettes representing nude female figures such as those of Brassempouy, Mentone, Pont-À-Lesse (Belgium), Predmost and Willendorf, near Krems. The Neandertal type appears to have died out and Aurignacian man belongs to the modern type represented at Combe Capelle. If the evidence of the figurines is to be accepted, a steatopygous race was at this time in existence, which Sollas is inclined to connect with the Bushmen[47].

The Solutrian stage is characterised by the abundance of the horse, replaced in the succeeding period by the reindeer. The Solutrians seem to have been a warlike steppe people who came from the east into western Europe. Their subsequent fate has not been elucidated. The culture appears to have had a limited range, only a few stations being found outside Dordogne and the neighbouring departments. The technique, as shown in the laurel-leaf and willow-leaf points, exhibits a perfection of workmanship unequalled in the Palaeolithic Age, and only excelled by late prehistoric knives of Egypt.

The rock shelter at La Madeleine has given its name to the closing epoch of the Palaeolithic Age. The flint industry shows distinct decadence, but the working in bone and horn was at its zenith; indeed, so marked is the contrast between this and the preceding stage that Breuil is convinced that "the first Magdalenians were not evolved from the Solutrians; they were new-comers in our region[48]." The typical implements are barbed harpoons in reindeer antler (later that of the stag), often decorated with engravings. Sculpture and engravings of animals in life-like attitudes are among the most remarkable records of the age, and the polychrome pictures in the caves of Altamira, "the Sistine chapel of Quaternary Art," are the admiration of the world[49].

In the cave of Mas-d'Azil, between the Magdalenian and Neolithic deposits occurs a stratum, termed Azilian, which, to some extent, bridges over the obscure transition between the Palaeolithic and Neolithic Ages. The reindeer has disappeared, and its place is taken by the stag. The realistic art of the Magdalenians is succeeded by a more geometric style. In flint working a return is made to Aurignacian methods, and a particular development of pygmy flints has received the name Tardenoisian[50].

The characteristic implement is still the harpoon, but it differs in shape from the Magdalenian implement, owing to the different structure of the material. Painted pebbles, marked with red and black lines, in some cases suggesting a script, have given rise to much controversy. Their meaning at present remains obscure[51].

The question of prehistoric chronology is a difficult one, and the more cautious authorities do not commit themselves to dates. Of late years, however, such researches as those of A. Penck and E. BrÜckner in the Alps[52] and of Baron de Geer and W. C. BrØgger in Sweden[53], have provided a sound basis for calculations. Penck recognises four periods of glaciation during the pleistocene period, which he has named after typical areas, the GÜnz, Mindel, Riss and WÜrm. He dates the WÜrm maximum at between 30,000 and 50,000 years ago and estimates the duration of the Riss-WÜrm interglacial period at about 100,000 years. According to his calculations the Chellean industry occurs in the Mindel-Riss, or even in the GÜnz-Mindel interval, but it is more commonly placed in the mild phase intervening before the last (WÜrm) glaciation, this latter corresponding with the cold Mousterian stage. At least four subsequent oscillations of climate have been recognised by Penck, the Achen, BÜhl, Gschnitz and Daun, and the correspondence of these with palaeolithic culture stages may be seen in the following table[54].