Snakes (Ophidia) are the newest and most flourishing branch on the reptilian family tree, whose trunk and lower limbs are dead or dying. They differ from lizards mainly in their elongated and limbless form (which, however, had been foreshadowed by certain lizards) and more particularly in the formation of the mouth. Instead of a solid union of the bones of the skull, many of the bones, especially about the mouth, are connected by an elastic ligament, allowing the snakes to open their mouths widely enough to swallow larger prey than otherwise would be possible. The palatal bones, as well as the jaws, bear small, solid, recurved and pointed teeth, replaced by others from the same root pulp when lost; they have little chewing power, but are useful to seize and hold food which is then slowly swallowed by the snake gradually working its jaws ahead and over the object, until the muscles of the throat can grip it and slowly work it downward into the tubelike stomach. Serpents strive to turn their prey and swallow it headfirst. The tongue in all serpents is a slender, extensible organ, forked at the tip, usually black, and always seen protruded and waving about when a snake is disturbed. Uninformed people call it a "stinger," but it is merely the animal's tongue and used as such. It serves the additional purpose of an instrument of The eyes, which may be rudimentary in the burrowing species, or large in those of nocturnal habits, have no eyelids, and are covered with a transparent film of skin that is sloughed off whenever the skin is shed, which happens frequently in young, growing individuals, but only annually in adults, as a rule; and for a day or two snakes are blinded by the loosening of this covering. No snake has ear openings, and their hearing is dull. The sense of smell, however, is well developed, and it is probable that these animals obtain much food by its aid, even following a trail by the nose. Serpents travel on their bellies, moving their bodies in lateral undulations, and often running with amazing swiftness. Every pair of ribs is connected at their lower ends with one of the large abdominal scales, or "scutes," and it is generally believed that the creature moves by the pressure and pushing of these scutes and rib points on the ground; but Boulenger, a leading authority, thinks that their importance has been somewhat exaggerated, although of undoubted use for the purpose of climbing, at which some species are remarkably adept. Some snakes lay eggs with a tough, parchmentlike shell; others retain them within the body until the young are fully developed. Snakes do not migrate nor wander far from their birthplace in search of food. Desert dwellers The order is divided into nine families, which will now be considered in the order arranged by G. A. Boulenger of the ZoÖlogical Society of London. The first four families are small, wormlike, burrowing creatures, with a large number of species distributed in warm countries throughout the world, and regarded as relics of an ancient type. The beautiful coral snake of South America, which grows to a yard in length and is only partly subterranean in habit, leads from these to the great family of boas and pythons (BoidÆ) which contains the biggest serpents that exist, or so far as we know, ever have existed. The members of this family have vestiges of pelvis and hind limbs, appearing externally as clawlike spurs. The BoidÆ comprise sixty or seventy species and the range of the family is world-wide. They mostly prefer wooded districts, climbing trees, assisted by the short and partly prehensile tail. Some are semiaquatic. All are rapacious, and feed by preference on warm-blooded creatures. The family is divided into two subfamilies, PythoninÆ and BoinÆ, but the difference between them is confined mainly to certain bones in the skull. The pythons belong entirely to the tropics of the A famous python is the six-foot, tree-dwelling carpet snake of Australia, black, beautifully marked with a pattern of yellow dots. A very large species is the reticulated python of Indo-China and the Malayan region, having an arrangement of dark lozenges on a lighter ground. India has a similar species, reaching a length of thirty feet, marked with reddish brown patches on a yellowish ground. This (Python molurus) is the one most often seen in zoÖlogical collections on account of its hardiness; but it is a savage creature, almost untamable. Like others of these big serpents it is able to make very long fasts; indeed their life, in this respect, seems to consist of gorges, followed by long periods—sometimes several months—of fasting and repose, entirely voluntary. It appears from observation of captive specimens that they have individual preferences for a certain kind of food, and perhaps wait for it; thus one in the Jardin des Plantes, Paris, refused various toothsome animals for months until a goose was offered, which it seized hungrily, and then sulked through long weeks until another goose was given. Africa has two pythons, one (P. regius) confined to West Africa, the other (P. sebÆ), common from the Sudan to the Cape. "The latter," William C. Scully says, "is the largest of African snakes, occasionally attaining a length of more than twenty feet, with a circumference of eighteen inches. One is recorded of twenty-five feet. It principally frequents rocky chasms in moist, warm forests. It is not dangerous to man, being quite nonvenomous, but it will fight fiercely if attacked, and the long, sharp, "So far as I know the python is the only snake which incubates its eggs. Such, numbering from thirty to fifty at a brood, and weighing about five and a half ounces each, are usually laid in a deep rock crevice or in the deserted burrow of an ant bear or hyena. The mother coils herself over and around them." Let us turn now to the boas. Popularly the whole tribe is frequently spoken of as "boa constrictors," but that is the scientific name of only one among several species, the Boa constrictor of the West Indies and tropical South America. It is the one most common and best known, and, as it is easily tamed, is the one often seen in the hands of performers with serpents in circuses, and exhibited in menageries. In many places in South America the natives, according to Leo Miller, keep them running at large about their huts to catch rats. In forested regions they spend most of their time in trees, but in an open country lie about on the ground, retreating when alarmed into some hole, as of a viscacha—their favorite prey on the plains. Far greater and much more dreaded by the natives is the great water boa, or anaconda (B. murinus), of the Amazonian region, which is the longest of American snakes, and the worst foe of such river-loving creatures as capybaras and iguanas. The color scheme of the anaconda is greenish yellow above, with a single, or two alternating series, of large, blackish transverse spots, and one or two lateral series of blackish eyespots with white centers. The lower parts are whitish, spotted with black. The anaconda is very aquatic, and is usually found submerged close to the banks of the river, on the lookout for its prey. Although mammals and crocodiles are occasionally eaten by this snake, it prefers birds, these being constricted and eaten under water. Only a single instance of an anaconda having attacked a man is on record. Although it grows to a length of over thirty feet, it is sexually mature when about half that length. Various very slender and agile species, the tree boas, belong to the tropical American forests, one of which is called the "rainbow" boa because of its marvelous iridescence in the sunlight. Another large species inhabits Central America and Mexico; and two small, brown secretive snakes, the "rubber" boas, more commonly known as "double-enders," because their blunt tails closely resemble their heads, are found in California and northward to British Columbia. The remainder of the family are scattered from Africa to the South Sea Islands. ORDINARY SNAKES (COLUBRIDÆ)We now come to the family ColubridÆ, which embraces nine-tenths of all the modern serpents of the world. The more hardy species are to be found The best arrangement of the ColubridÆ is that by Boulenger, who, adopting DumÉril's terms, has divided them into three series according to the character of the teeth. 1. Aglypha—All the teeth solid and not grooved. Harmless, that is, not venomous. 2. Opisthoglypha—One or more of the posterior maxillary teeth grooved. Mostly poisonous; a few tropical species. 3. Proteroglypha—Anterior maxillary teeth grooved or tubular. Deadly poisonous; cobras, coral snakes, etc. The immense family ColubridÆ is divided into several subfamilies, the first and most extensive of which is the ColubrinÆ, in which are associated all the "harmless" snakes in the world except the boas and pythons. None exceeds twelve feet long, and most of them are much smaller. Nearly all lay eggs, but some bring forth large broods of living young, among which are our water snakes, and the striped "garter" and "ribbon" snakes so numerous in our meadows and gardens. These striped snakes (Eutainia) exist in a great number of "species" or varieties most confusingly varied in coloring, some having no stripes whatever. They are very hardy, living far toward the north in Canada, and are the last to go into hibernation and the first to reappear in spring. For this winter sleep they burrow deeply into soft soil, or where rocky places exist, seek deep crevices. The water snakes of the genus Tropidonotus follow, with many representatives in all temperate countries, one of which is the "common grass Various small, ground-keeping snakes lead to another conspicuous American group, the racers and black snakes of the genera Spilotes and Zamenis, of which species and near relatives are numerous in Europe and Asia, a Malayan example growing to a length of ten or more feet—probably the longest of colubrines. Three different "black snakes" are known among us. The largest is the The genus Coluber, to which belongs the famous Æsculap snake of central Europe, is represented among us by a series of large and gayly colored species. One is the yellowish, brown-blotched fox snake of the prairie States, which is a ground keeper and a great hunter. In search of rats and mice it often haunts haystacks and barns where it should be welcome. "One snake is worth a dozen traps, for the reptile prowls into the burrows and nests of rats and mice and eats the entire brood." Similar in size (six feet) and habits is the brilliant red-and-crimson corn snake of the Southern States, which is a great mouser and also an agile climber after nests of birds, whose eggs and young it likes. Another, even larger, coluber of the South is the four-striped chicken snake, useful in its pursuit of small rodents, but, like the pilot black snake, with too great a fondness for hen's eggs and young poultry to be liked by farmers. The big, gray, blustering "bull snakes" of the southern and western parts of the Union take their name from their habit of emitting a loud and prolonged hiss when annoyed. They keep on, and under, the ground in sandy regions, feed on small mammals and birds, and are powerful constrictors; they are also noted for morose and savage dispositions. Next to these repulsive reptiles come in classification Passing over a number of small, smooth-scaled serpents, of which the pretty ringneck is an example, we come to the important genus Ophiobolus, which contains the king snakes, milk or house snakes, coral snakes and others, represented in the Old World by the genus Coronella. They vary in size from fourteen inches to six feet, and in color from gray with dark blotches to a ringed pattern of red, black, and yellow, often of brilliant beauty; but there is much individual variation. The king snake might easily furnish material for a long chapter. Its name follows from its known disposition to pick a quarrel and fight with any serpent it meets, big or little; and quite independent of whether it is hungry, for it is as fond of eating its own kind as it is of lizards, toads, mice, birds and anything else that comes in its way on the ground, for it is not much of a climber. Our books are full of incidents of its destruction of poisonous species, and the popular belief is that it hunts for, and relentlessly pursues rattlesnakes, copperheads, etc., but the authorities assure us this is not so. If it accidentally encounters a rattler or moccasin, it kills, and perhaps eats it; but it does the same with any other serpent. It is an exceedingly quick and powerful constrictor, and careless of bites, for it is The common northern representative of the genus is the house snake or milk snake—names given to several other species; it is also known as "checkered adder," because of the general resemblance of its blotched form to the dreaded copperhead. It is gray above, with a series of large, chestnut-brown saddles on the back, smaller blotches alternating with them along the sides; the belly is white, marked boldly with square black blotches. The pattern and tints vary widely. This snake is a lover of warmth and a hunter of mice and rats, wild and domestic; and in search of them it frequents pastures and damp meadows, where such wild game abounds, comes much about stables and houses, and often creeps into the rural dairies that are usually close to springs. Serpents with these inquisitive habits are familiar in all parts of the world, and from time immemorial have been accused, among other iniquities, of milking cows and goats, and of drinking and spoiling milk and cream on the shelves in dairies and cellars. These beliefs survive among country people to this day, as I found out a few years ago by an extensive correspondence of inquiry, in which We will mention only one other sort of our harmless colubrines—the "hognose," "puffing adder," "spreading adder," as it is variously known; but the name hognose is the best. Its genus is Heterodon. Two species are common all over the eastern half of the United States and Canada, one an ugly mottled gray, the other black. They are about two feet in length, thick-bodied, with roughly keeled scales, a flat head and a pointed, upturned snout—altogether very unhandsome and forbidding-looking reptiles; and they profit by this in an attempt to frighten away whatever alarms them, while in reality themselves almost (sometimes quite) paralyzed by fear. CORAL SNAKES, COBRAS, AND SEA SNAKESThe flattening of the head and neck practiced by the hognose as a gesture of readiness to fight, whether true or false in its implication of ability, is found among several non-poisonous colubrids elsewhere and indicates their approach in kinship to the "hooded" cobras that are the foremost representatives of the venom-bearing members of the ColubridÆ. It will be recalled that we have been The principal tooth-bearing bone in a serpent's mouth is the forward half of the upper jaw, termed the maxillary. The maxillary of each side is connected with its fellow by a small, single bone in front (the premaxillary) and otherwise is connected with the loosely connected bones of the skull by those elastic cartilages that enable the mouth to expand and take in prey of a size more than equal to the snake's head when the mouth is shut. In the serpents that do not possess a poisoning apparatus the teeth on the maxillaries are alike in size, and solid; but in the venomous kinds some of the teeth are enlarged and grooved or channeled to conduct a flow of poison into the wound made by biting. This is the case with the poison-bearing sections of the ColubridÆ mentioned above, and their difference is in the relative position of the poison-conducting teeth or "fangs" on the maxillaries. In the Opisthoglyphs these teeth are situated near the posterior end of the maxillary, and are grooved on the rear side, where they receive the poisonous fluid from a sac in the cheek. The greater number of species of this group are residents of the Old World, although we have several representatives along our Mexican border, and more southward, especially in the tropics. Most of them are little dangerous to mankind, as it is difficult for them to inflict a wound by "striking." They first seize their prey and then use their rather short fangs. The poison has a paralyzing effect, In the Proteroglypha, on the contrary, the poisoning teeth, in all cases small, are situated near the front of the maxillaries, and they are much more dangerous reptiles, for they include the coral snakes and cobras (ElapinÆ) and the sea snakes (HomalopsinÆ), which are able to strike their teeth into anything they successfully attack. The coral snakes (genus Elaps) derive their name from the broad bands of coral-red that encircle their bodies in most species, with narrow rings of black and yellow between. These brilliant colors, combined with the luster of the smooth scales, make them among the most beautiful of serpents, and a common species of our Southern States is called the harlequin. The genus is exclusively American, and nearly all belong to the tropics, where the largest become five feet long, and their bite is deadly to man. They keep to the ground, and much of the time under it, and are cannibalistic in their diet. The body is slender and cylindrical, the head small, and the eyes like beads. They are indocile, quick-tempered, and very dangerous to handle, despite the fact that they do not always resist being disturbed. Hence the widely prevalent opinion that they are harmless is a perilous delusion fostered by the fact that certain The remainder of the elapine serpents (about 125 species) belong to Africa and the Orient. Typical of them are the cobras of the genus Naja, of which the species (Naja tripudians) met with from Turkestan to southern China and the Malay islands, and named by Portuguese explorers "cobra de capello" (hooded snake), is world-famous. Several species inhabit Africa and differ little from the Indian cobras, but are equally deadly. The fangs in all this group are small and are fixed in the extreme front of the upper jaw, not being erectile like the long fangs of the rattlesnakes and vipers. Cobras vary much in coloration, and Mr. Scully reports that he has killed South African specimens of light yellow, jet black and all intermediate hues. The cobra is a fierce fighter and, when reared up, with expanded hood, looks very formidable. Anterior to the head the ribs lengthen and then gradually shorten to normal dimensions. These lengthened ribs, about twenty in number, lie, when the snake is quiescent, more or less laterally along the spine. But when the snake becomes excited, the The cobra is found all over South Africa, but is especially plentiful in the dry, sandy deserts northwest of the Cape. There extensive colonies of large mice abound, patches of ground being thickly honeycombed with burrows. In these the cobras dwell—apparently, as in the case of the puff adders, on the best of terms with their hosts, upon whom they principally feed, reminding one of the tenancy by the Western rattlesnake of prairie-dog "towns." A close relative of the cobra is the ringhals (i. e., ringneck), known as the "spitting snake," the explanation of which Mr. Scully furnishes from personal experience thus: "The ringhals, when excited, exudes a quantity of venom, which drips down the fangs and lodges behind the abrupt, horny, lower lip. Upon this the angry snake directs a blast of air through its extensible windpipe, with the effect that a jet of fine venom spray is emitted toward an enemy. This jet may reach a height of six feet. That the eyes are aimed at I have proved by experiment. If the poison reaches them blindness, which may be permanent, results. The bite of the ringhals is highly venomous, but the snake appears to prefer disabling its enemy by means of the spray of venom." The most novel and interesting of Mr. Scully's contributions to African herpetology, however, is his story of the mamba (Dendraspis angusticeps), which he calls "the head of the family." It is the longest venomous snake in the world, probably "The mamba has the habit of lying coiled among the branches adjacent to a footpath in a forest. Woe to the passing wayfarer in such a case! If he touch a twig, and thus impart the least tremor to the snake's lair, a lightning-swift stroke upon face, neck, or arm seals his doom. Such a stroke may be delivered either forward or sideways, with equal speed and effectiveness." The most feared of the cobra tribe in India and eastward is the king cobra, or hamadryad, which often exceeds a dozen feet in length and is "the largest, boldest and most dangerous of all venomous snakes," in Boulenger's opinion, "for when disturbed it does not content itself with merely sitting up and expanding its hood, but will almost invariably attack." Fortunately it is not numerous anywhere in its range from the Himalayas to the Far East; and it is useful in that it feeds exclusively on snakes, small pythons, kraits, rat snakes, Nearly all the serpents of Australia belong to ElapinÆ, and are exceedingly dangerous. Among them are the "black snakes," the females of which are called "brown adders"; the "tiger," so called from its colors; and most dreaded of all, the "death adder," which is distinguished by a peculiar tail end, and by the fact that the head is made distinct from the body by a narrow neck, giving it a viperlike appearance. When disturbed it flattens out the whole body. A few words about the sea snakes will close our account of the poison-bearing colubrids. These are set apart in the subfamily HomalopsinÆ, on account of the structure of the tail, which is flattened vertically into a combination of swimming organ and rudder, for they live in the estuaries of Oriental rivers, and go far out to sea in their search for food; and are to be met with from the Persian Gulf to Polynesia and Japan. All are very poisonous, feed mainly on fishes and produce living young; and all are clothed in varied and brilliant colors. Living in the sea, or in tidal inlets, their movements in the clear blue water are agile and elegant; and in the Bay of Bengal they are sometimes seen congregating in large shoals. We turn now to the last and most advanced family of serpents, the vipers, rattlesnakes, moccasins, copperheads, and so forth (ViperidÆ). VIPERS, MOCCASINS AND RATTLESNAKES"Viper" is an old French-Latin word, meaning The original little "viper" of Europe and Asia is more a nuisance than a peril, for it is rarely more than a foot long, and its bite would be fatal only to a small child. A larger species, the sand viper, ranges from Italy to Armenia. India, Burma and Siam, however, have a member of this group which is pronounced by Sir J. Fayrer as next to the cobra the most dangerous serpent of the East—the daboia, or Russell's viper. It is nocturnal, not aggressive, and makes a loud hissing when anyone comes near it, so that it does not cause as many human deaths as it might; but frequently kills grazing cattle by biting them on the nose. The greatest and worst of these snakes belong to Africa, where the northern deserts are infested with two greatly dreaded species—the horned and the saw vipers. The former has two sharp hornlike protuberances above the eyes, and Canon Tristram writes that its usual habit is to coil itself on the sand, where it basks in the impress of a camel's footmark, and thence suddenly to dart out on any passing animal. Horses as well as men are in constant terror of it, for it will attack without any provocation. The worst of the African vipers, nevertheless, is the puff adder, which ranges over the whole continent, and may grow to a length of six feet, with a girth equal to a man's thigh. "The coloration of the puff adder," Mr. Scully writes, "is in groundwork a series of delicate All the pit vipers are American except a few species in southern Asia, some of which are arboreal in habit and have red prehensile tails. Our American species fall into two genera: Ancistrodon, the moccasins (no rattles), and Crotalus, the rattlesnakes. The "upland moccasin" of the South is the "pilot" or "copperhead" of the North, where it still exists in forested and rocky districts from Connecticut and the Great Lakes to Texas, and is particularly abundant in the rough hills beside the Hudson River, and thence southward along the Alleghenies. Its general hue is yellowish brown, becoming chestnut or coppery red on the head and end of the tail, which terminates in a hard point. Along the back, meeting irregularly on the midline, are chestnut-hued blotches that divide on the sides, forming inverted Y's; the belly is yellowish with distinct black blotches, leaving the throat clear. After one has seen a copperhead he is not likely to confuse it with the milk snake or any other. Its haunts and habits are much the same as those of the eastern rattlesnakes, nor do I consider it any more aggressive in spite of a rather over-blackened reputation, nor so deadly in the effects of its weaker venom. It is bad enough, however, and should be killed on sight wherever children or pet dogs are likely to meet with it. This upland moccasin is named in science Ancistrodon contortrix; its brother species, the water moccasin, is A. piscivorus. The moccasin is a larger, heavier snake than the copperhead, and a dweller in the sluggish rivers and swamps of the Gulf States and northward to North Carolina and Kentucky. The moccasins commonly lie on the branches of bushes at the edge of the water; and if escape from danger be possible they quickly drop into the water and swim away beneath it to some hiding place. If suddenly surprised they coil and open the mouth widely toward the intruder, showing its white interior that has given them the name "cotton mouth" among the Closely allied to the copperhead and moccasins are two very dreadful snakes of the American tropics—the "fer-de-lance" of the French islands of the West Indies, and the "bushmaster" of Brazil. The former reaches a length of six feet, and the bushmaster, or surukuku, as the Indians name it, to twice that length, thus rivaling the great viper of India. Both have all the ferocity and power of their race exaggerated to the limit, and hundreds of human lives are sacrificed to them every year. Every traveler has thrilling tales about them. Leo Miller, a cool-headed man of science, takes very seriously the fear this creature inspires. He reminds us that a bushmaster ten feet long has fangs an inch and a half long, and injects nearly a tablespoonful of poison at a single thrust. A man would survive such a dose but a few minutes. When once a bushmaster fell from a tree branch into his canoe everybody in it sprang overboard, and some narrowly escaped drowning. Such deadly creatures would make the tropical world unendurable were it not that most of the time they are sluggish and peaceful; but a little fright, or a protective instinct in regard to their eggs, sets them off with the suddenness of a released spring. In taking up the rattlesnakes we have a sure guide in Dr. Leonhard Stejneger's "Report," describing all the species of North America (the group CrotalinÆ is confined to this continent, Central America, and a single species in South America). The special peculiarity of the group is the queer "rattle" (crotalus) at the end of the tail. This consists of a series of loosely connected, somewhat cone-shaped, horny The smallest of the crotalids are the ground rattlers (genus Sistrurus), of which we have two species, and there is one in Mexico. The northern kind, widely known by its Indian name "massasauga," ranges from eastern Pennsylvania and Ontario to northern Minnesota and Kansas, and thence to Texas. The Southern States have a second species commonly called "ground rattler." Both are grayish brown with chestnut or darker dorsal blotches, and are inhabitants of the prairies, with their swamps and marshes. The largest do not exceed forty inches, and their bite is correspondingly weak in effect. The remainder of the rattlesnakes belong to the genus Crotalus. The commonly seen species of the region east of the dry plains was named Crotalus horridus by LinnÆus, and this is one of the few instances in which his name has defied change by the systemists. It Far more formidable than this is the diamond-back (C. adamanteus) of the low, coastal region from North Carolina to the lower Mississippi River and throughout Florida, where it is far more common than is desirable. This rattler may exceed eight feet in length, and has corresponding power of harm. It is partial to the neighborhood of water, where its ground-running prey is most numerous; hence it is frequently spoken of as the "water rattlesnake," to distinguish it from the banded species, or "timber rattlesnake" of the same region, which is more habituated to forested districts, with rocks. A race of C. horridus, usually large and vicious, exists in the coast swamps, and is locally called the "canebrake rattler." The diamond back itself takes its name from the lozenge-shaped patches of dark color formed on its upper surface by the crossing of diagonal narrow bands of bright yellow on a greenish gray ground. The literature relating to this terrifying snake would fill hundreds of pages. Raymond L. Ditmars of the New York ZoÖlogical Park, gives this description: "Most deadly of the North American poisonous snakes, and ranking in size with the largest of the A large and very showy western analogue of the diamond-back, known by the sinister specific name atrox, occurs from central Texas to California. One of its varieties is red, with darker red markings and a white tail. The familiar rattlesnake of the plains east of the Rocky Mountains is Crotalus confluentus, which is of moderate size and dull hue; its mainstay of food is found among the prairie-dog towns. A similar but smaller species (C. oregonus) takes its place west of the Rockies, from British Columbia to southern California. The "tiger" rattlesnake, yellow barred with black; the "horned" rattler, which, like the Egyptian horned viper, has a trick of advancing sidewise, and consequently has the popular name "sidewinder"; and the slender green rattlesnake, are small species of the deserts along the Mexican border. |