MATERIALS, ACKNOWLEDGMENTS AND METHODS

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At a late stage in the preparation of this manuscript a total of 5,457 specimens had been examined. For the most part these were conventional study-specimens; that is to say, they were stuffed skins with the skulls separate and each was accompanied by the customary data as to locality of capture, date of capture, name of collector, external measurements and sex recorded on the labels by the collectors. Skulls unaccompanied by skins, nevertheless, comprised a large share of the total and a small proportion was made up of skins unaccompanied by skulls, mounted specimens, skeletons, and entire animals preserved in liquid.

It was the recognition of this need for specimens from extensive areas from which no specimens previously had been collected that influenced me, approximately a year after the study was begun, to allot for it a long span of time. The procedure adopted, in general, was to study the weasels of one species from a given geographic area in so far as the material warranted, then lay this aside until additional critical material could be obtained, and finally, some months or a year later, complete the account. In this fashion the manuscript of the American weasels received my attention in each of the past twenty-five years (September, 1926 to date of publication). This is a confession of fact rather than a recommendation of procedure. This type of procedure unduly delays the diffusion of knowledge and for a variety of reasons justifiably annoys other students of the subject. Nevertheless, many gaps have been filled that otherwise would have remained open. Although specimens to solve several problems still remain to be collected and studied, it seems that a point of diminishing returns has now been reached, which, in fairness to all concerned, calls for publication of the results so far obtained.

For assistance in the entire undertaking, I am more indebted to Miss Annie M. Alexander than to any other one person; she provided the means by which specimens from critical areas were obtained, made it possible to examine the European collections, and assisted in other ways. The late Professor Joseph Grinnell and Mr. Charles D. Bunker, among others, gave truly valuable encouragement and assistance.

Collections containing weasels which were examined in the study here reported upon were as follows:

  • Acad. Nat. Sciences of Philadelphia
  • American Mus. Nat. History
  • Baylor University
  • Berlin Zoological Museum
  • Boston Society of Natural History
  • Brigham Young University
  • British Museum of Natural History
  • California Academy of Sciences
  • Carnegie Museum
  • Charleston Museum
  • Coe College
  • Collection of J. Arnold
  • Collection of Stanley C. Arthur
  • Collection of Rollin H. Baker
  • Collection of William Bebb
  • Collection of R. H. Coleman
  • Collection of Ian McTaggart-Cowan
  • Collection of Stuart Criddle
  • Collection of John Cushing
  • Collection of Walter W. Dalquest
  • Collection of William B. Davis
  • Collection of J. M. Edson
  • Collection of Ralph Ellis
  • Collection of John Fitzgerald, Jr.
  • Collection of Mr. Green
  • Collection of Ross Hardy
  • Collection of Donald V. Hemphill
  • Collection of L. M. Huey
  • Collection of R. W. Jackson
  • Collection of Stanley G. Jewett
  • Collection of E. J. Koestner
  • Collection of J. E. Law
  • Collection of A. H. Miller
  • Collection of Lloye H. Miller
  • Collection of R. D. Moore
  • Collection of J. A. Munro
  • Collection of O. J. Murie
  • Collection of Robert T. Orr
  • Collection of Arthur Peake
  • Collection of Kenneth Racey
  • Collection of William B. Richardson
  • Collection Rocky Mt. Spotted Fever Lab.
  • Collection of Victor B. Scheffer
  • Collection of William T. Shaw
  • Collection of O. P. Silliman
  • Collection of W. E. Snyder
  • Collection of Frank Stephens
  • Collection of T. C. Stephens
  • Collection of D. D. Stone
  • Collection of Myron H. Swenk
  • Collection of Joe and Dean Thiriot
  • Collection of John Tyler
  • Collection of Jack C vonBloeker
  • Collection of Alex Walker
  • Collection of Edward R. Warren
  • Colorado Museum of Natural History
  • Charles R. Conner Museum
  • Cornell University
  • Donald R. Dickey Collection
  • Field Museum of Natural History
  • Florida State Museum
  • Fresno State Junior College
  • Humboldt State Teachers College
  • Illinois Natural History Survey
  • Iowa State College
  • Iowa Wesleyan College
  • Kansas State Agric. College
  • Leland Stanford Junior University
  • Leningrad Academy of Science
  • Los Angeles Mus. Hist. Art and Sci.
  • Louisiana State University
  • Mt. Rainier Nat'l Park Collection
  • Museum of Comparative ZoÖlogy
  • Mus. Polonais d'Hist. Nat., Warsaw
  • Mus. Vert. ZoÖl., Univ. California
  • Museum of ZoÖlogy, Univ. Michigan
  • National Museum of Canada
  • Naturhistoriska Ricksmuseum, Sweden
  • Neuchatel University Museum
  • New York State Museum
  • Ohio State Museum
  • Oklahoma Agric. and Mech. College
  • Ottawa University, Kansas
  • Paris Museum
  • Provincial Museum of British Columbia
  • Royal Ontario Museum of ZoÖlogy
  • San Diego Society of Natural History
  • State Hist. and Nat. Hist. Soc. Colo.
  • State Normal School, Cheney, Wash.
  • Texas Cooperative Research Collection
  • United States National Museum
  • University of Arkansas
  • Univ. California Mus. Palaeo.
  • University of Idaho
  • Univ. Kansas Mus. Nat. History
  • University of Minnesota
  • University of Notre Dame
  • University of Oklahoma
  • University of Oregon
  • University of South Dakota
  • University of Utah
  • Univ. Washington Museum of ZoÖlogy
  • University of Wisconsin
  • Univ. Zool. Mus., Copenhagen

The largest single collection is in the United States National Museum, where the specimens of the National Museum proper and the United States Biological Surveys Collection, together, provide essential materials including a large share of the holotypes. Specimens in all of the North American collections including Canada and MÉxico have been made available, by loan, and in 1937 materials were examined in the principal collections of northern and central Europe. After the materials in North American collections were assembled, special effort, with considerable success, was made in each of several winters, to obtain specimens from areas not previously represented in collections.

To the many persons who were in charge of the collections consulted, to those who at my request sought critical specimens, and to those who assisted in various stages of assembling data and in preparation of the manuscript, I am grateful indeed. Likewise, I am deeply appreciative of the grants-in-aid received from the Carnegie Institution of Washington, the University of California Chapter of Sigma Xi, the John Simon Guggenheim Memorial Foundation and the Kansas University Endowment Association. I am mindful also of an obligation to those who appropriated funds, by legislative action, for research use by The University of California and The University of Kansas.

For assistance with the illustrations I am indebted to the late Major Allan Brooks for Plate 1 , to Mrs. Mary Blos for figures 25-31, to Miss Ann Murray for figures 11-13, to Mr. W. C. Matthews for all the photographs, to Mrs. Freda L. Abernathy for figures 2-9, 18-22, 24, and for retouching all the photographs except the following which were retouched by Mrs. Virginia Unruh: figs. d of plates 2, 3, 4, 9, 10, 11, 16, 17; figs. i of plates 5, 6, 7; figs. h, j, k of plate 7; figs. f and g of plates 12 and 13; and figs. c and d of plate 14. To Mrs. Unruh I am further indebted for figures 1, 16, 17 and 23 and for much terminal assistance with preparing most of the illustrations for the engraver.

The methods of study, after specimens were assembled, included first comparisons of specimens of like age and sex from each of several localities to ascertain the constant features by which full species were distinguishable, one from the other. For example, it was found that in every individual from Trout Lake, Washington, of the species here designated Mustela erminea, the postglenoidal length of the skull amounted to more than 47 per cent of the condylobasal length whereas it was less than 47 per cent in all individuals here designated as Mustela frenata, from the same locality. Testing of specimens from other localities by means of this and other selected characters permitted the outlining of the geographic ranges of the full "species-groups." By comparing specimens of other nominal species and by examining specimens from localities geographically intermediate between the nominal species, I found intergradation and therefore arranged the nominal species as subspecies of a single species. Intergradation here is understood to be the result of crossbreeding in nature between two kinds of animals in the area where the geographic ranges of the two kinds meet. Presence of intergradation between two kinds of weasels was basis for according them subspecific rank. Absence of intergradation in nature at every place where the geographic ranges of two kinds met or overlapped, and absence of intergradation by way of some other kind, or chain of kinds, was basis for according each of the two kinds full specific rank. By thus applying the test of intergradation, or lack of it, I found that there were four full species of weasels, of the subgenus Mustela, in all of the Americas.

Next, the specimens of one species were arranged in trays in a geographic sequence. The specimens from any one locality were segregated by sex and under one sex from one place were arranged from oldest to youngest, that is to say by age. The four series with the largest numbers of individuals of a given age were selected. Seventeen cranial measurements and three external measurements were recorded for each individual of each of these four series. For each measurement, the coefficient of variation, standard deviation and probable error were computed. The four samples subjected to such analysis were a series of adult males, one of adult females, one of subadult males and one of subadult females. Also, studies of each sex were made to ascertain seasonal changes in pelage. After data were obtained on ontogenetic (age) variation, secondary sexual variation, seasonal variation, and degree of individual variation by studying specimens in the manner described above, tests were made for subspecific (geographic) variation by comparing series of specimens of like sex, age and season, from different localities. For each one of several geographically variable features noted, a map was prepared for animals of each sex. When all the data thus obtained were codified, subspecific ranges were, in a sense automatically, obtained. On the resulting map showing geographic ranges of subspecies for a species, a type locality was accurately plotted for each name that had been applied to the species, and names then were applied in accordance with the international rules of zoÖlogical nomenclature.


                                                                                                                                                                                                                                                                                                           

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