MYRMELEON LIBELLULOIDES.Plate XLVI. fig. 1. Order: Neuroptera. Section: Filicornes. Family: MyrmeleonidÆ, Leach. Genus. Myrmeleon, Linn. &c. Myrmeleon Libelluloides. Alis fusco punctatis maculatisque corpore nigro flavoque maculato. (Expans. Alar. 4 unc. 9 lin.) Syn. Myrmeleon Libelluloides, Linn. Syst. Nat. 1. 2. p. 913. 1. Fabr. Ent. Syst. 2. 92. 1.? Latr. Gen. Crust. & Ins. 3. 191. Libella turcica major alis locustÆ, Pet. Gaz. 6. t. 3. f. 1. Habitat: Smyrna (Drury). "In Europa australiori et per totam Africam" (Latreille). Head dark yellow, divided by a black line, which runs along it from the thorax. AntennÆ black and clubbed, small at the base, and gradually increasing to their extremities. Thorax brownish yellow; having a black line running along its upper part, and two small black spots at the base of the superior wings, being covered with greyish hairs. Abdomen about an inch and a quarter long, yellow, with a black line on the top, and one on each side; the male having two small horny tails issuing from the extremity. Wings membranaceous and pellucid (the inferior ones being as long as the superior), and elegantly adorned with a great number of dark spots of various shapes and sizes. Fabricius gives the Cape of Good Hope as the habitat of this species, referring not only to the present figure, but also to that given in Vol. 3. Plate 41. which is said by Drury to have been brought from Sierra Leone. This is the more inexcusable, because Drury expressly observed in a note, "There is a species found near the Cape of Good Hope very much like this, but distinctly different," although in the synoptical appendix to the third volume, he gives the large species from Sierra Leone as a variety of M. Libelluloides. Plate XLVI. fig. 2. Order: Neuroptera. Section: Filicornes. Family: MyrmeleonidÆ, Leach. Genus. Euptilon, Westw. Gen. Nov. Hemerobius, Linn. Drury. Euptilon Ornatum. Viride, thoracis puncto antico abdomineque line dorsali nigris, alis hyalinis venis numerosissimis, strigisque duabus obliquis obscuris. (Expans. Alar. circ. 3 unc.) Syn. Hemerobius ornatus, Drury, App. vol. 2. Habitat: Dinwiddie, in Virginia. Head dark green. AntennÆ pectinated or combed. Eyes black. Thorax dark green, with a black patch next the head. Abdomen dark green, with small rings of yellow, and a small black line running along the upper side from the thorax to the extremity. Wings membranaceous and pellucid, or transparent, of equal length; the superior ones having two small transverse black stripes placed near their posterior edges, at about half an inch distance from each other. Latreille, in his Genera Crustaceorum et Insectorum, Vol. 3. p. 199, after describing the genus Chauliodes belonging to the family HemerobiidÆ, observes, "Congenerica videntur insecta a Dom. Drury iconibus vivis expressa, tom. 1. Pl. 46. Fig. 2. 3." I have never seen the present species, but from the admirable accuracy of Moses Harris (by whom the figures were drawn) it is evident that the neuration of the wings of this species is identical with that of Myrmeleon, agreeing especially with the insect represented at Fig. 4. of this plate, and exhibiting the irregular longitudinal nerve below the subcostal nerve and the strong furcate nerve running along the middle of the wing; whilst, at the same time, the neuration is quite unlike that of Fig. 3. Hence I have no hesitation in placing this insect in the family MyrmeleonidÆ. It however disagrees with the genera of which that family is composed, viz. Myrmeleon, Linn. (Myrmecoleon, Burm.) Ascalaphus, Fabr. and Nymphes, Leach, in having bipectinated antennÆ, so that I have been compelled to establish a new genus for its reception. Can it be possible that, according to the not uncommon practice of the time, the specimen had been mended, and that pectinated antennÆ had been substituted in the stead of the ordinary ones of a Myrmeleon? CHAULIODES VIRGINIENSIS.Plate XLVI. fig. 3. Order: Neuroptera. Section: Filicornes. Family: HemerobiidÆ, Leach. Genus. Chauliodes, Latreille. Hemerobius, Linn. Drury. Chauliodes Virginiensis. Capite thoraceque nigris fulvo punctatis, alis latis hyalinis venis nigro punctatis. (Expans. Alar. 3 unc.) Syn. Hemerobius Virginiensis, Drury, App. vol. 2. Hemerobius pectinicornis, Linn. Syst. Nat. 1. 2. 911. 1.? Pal. Beauv. Ins. Afr. et Am. Neur. pl. 1. f. 2.? Habitat: Virginia. Head black, with a yellowish spot in the front, and another on the top. Eyes greyish, and shining like polished bell-metal. AntennÆ pectinated or combed, and longer than the thorax. Ocelli three. Thorax black, with three small yellowish spots on the posterior part. Abdomen almost black at top, Drury has figured the wings as of an uniform colourless appearance, which induces me to consider that it may be distinct from the LinnÆan H. pectinicornis, that species being described by the great Swede with "signaturis nervisque fuscis albo subarticulatis." From the paleness of the present species it is evident that the transverse nerves connecting the longitudinal ribs of the wings in the typical species of this genus have been overlooked, especially as the longitudinal nerves are correctly represented. This genus and Corydalis, seem to form a connecting link between Hemerobius and Perla. MYRMELEON AMERICANUM.Plate XLVI. fig. 4. Order: Neuroptera. Section: Filicornes. Family: MyrmeleonidÆ, Leach. Genus. Myrmeleon, Linn, &c. Myrmeleon Americanum. Thorace griseo nigro notato, alis hyalinis fusco et luteo punctatis, punctis nonnullis lineam centralem longitudinalem formantibus, abdomine fusco. (Expans. Alar. 5 unc.) Syn. Myrmeleon americanus, Drury, App. vol. 2. Habitat: New York. Head black, front grey and hairy. Mouth with four long palpi. Eyes dark brown, almost black. AntennÆ black, and as long as the thorax; being small at the base, and increasing in size to their extremities. Neck yellowish, striped with black, and covered with longish grey hairs. Thorax yellowish, with black stripes, and covered with long grey hairs. Abdomen, which is full an inch and a half long, brownish-coloured, and darker on the sides; being furnished at the extremity with two short hairy tails, seemingly of a horny substance. Wings of equal length, membranaceous and transparent, having a great number of small spots thereon of a dark brown colour; the superior ones being most spotted. Legs black, and covered with grey hairs; each of them being furnished at the tips of the tibiÆ with two long spines that bend inwardly towards each other. All the insects represented in these two plates belong to the genus Libellula of LinnÆus, or the family of Dragon-flies, LibellulidÆ. When the original edition of this work was published, there was no English work in which the natural history and curious transformations of this tribe of insects were detailed. Our author, therefore, in order to supply the deficiency, published the following interesting series of observations upon the subject:— "As I have not met with any English author who has given the natural history of the insects delineated in these two plates, I shall make no apology for its introduction; "It is not easy to determine whether they should be ranked among the water insects, or those of the land, nor shall I attempt here to ascertain it; my present business being only to relate the several circumstances attending them during their respective states in which they are passing from the egg to the complete animal: and although these observations have been confined to our English ones, yet they so exactly agree and coincide with those of foreign countries, (as my correspondents have assured me), that their nature and behaviour appear to be just the same; so that what is observable in ours, is at the same time applicable to the whole genus wherever found. "If we take a cursory view of the different ranks of animals that inhabit our globe, we shall hardly find one that can excite our wonder and astonishment more than this genus; nor is it from that general ignorance of the insect world, that reigns so strongly in these kingdoms, that I am emboldened to say this; but if we reflect that the beasts, birds, and reptiles are furnished with powers for living only in the air, and that even the amphibious tribes can perform the office of respiration only in that element: if we also consider that fishes, on the contrary, are unable to respire but in water, and when deprived of that must certainly perish, we cannot but conclude that all these animals are most wisely fitted with means and faculties for filling up the respective orders and ranks wherein they are placed. But let us cast our eyes on the subjects I am about to describe, and there behold a tribe of beings, who, as soon as they leave their eggs, subsist for a certain number of months, (I had almost said years), creeping and swimming in the liquid element; are there invested with organs and powers for existing and weathering out the utmost severity and intemperance of the winter; that afterwards as the spring and summer advances, and the period arrives when they are to appear in other forms, in the space of about half an hour those very organs and powers that before enabled them to live under water should be so entirely altered, the very natures and abilities of the creatures so changed as to permit them to quit their former element and place of residence, insomuch that all the remaining part of their lives is spent in the open air, furnished with wings, and flying about in the full glee of wanton liberty; that in a very few weeks after, having performed the business of generation, the same animals should die of mere old age, with their wings quite ragged and worn out, their strength exhausted, and all the powers of their bodies lost by a total imbecility and weakness, which but a little before enabled them to transport themselves through the air with the swiftness of a bird. If, I say, we reflect on all these circumstances, we cannot but allow them to be objects of a very extraordinary nature, and well adapted for leading the mind to the contemplation of their supreme Author, who has thought proper to exhibit to us these kind of insects, thus differing from almost all the animals in the creation. "They have been variously named by different authors, owing perhaps to the time when they wrote, or the progress natural history had made in the world. Some have called them by the name of Dragon Fly, others Adder Bolt, Balance Fly, Perla, Libellula or Libella. I shall prefer the last, as conveying an idea well known to English adepts. The caterpillars of them all live in ponds and stagnant waters (that are undisturbed by cattle) during the greatest part of their lives, and make their appearance under three general forms. The first is shewn by Fig. 1. 1.; the second by Fig. 2.; and the third by Fig. 3. See Plate 47. As these are the shapes in which they all appear, it will not be improper to mention each particularly. "The two caterpillars at Fig. 1. 1. Plate 47, belong only to or produce those that sit at rest, with their wings erect, (see Plate 48. Fig. 2.), and differ from both the others not only in size but in the roundness and slenderness of their bodies; at the extremity whereof they are furnished with three tails, each of which upon examination being found to be a kind of feather, and is an appendage that neither of the others have. Their behaviour also in the water is different from the others, being enabled to twist and bend their bodies in a more circular manner. [Subfamily Agrionides.] "The second sort of caterpillars, at Plate 47. Fig. 2. are much shorter and thicker in their bodies than either of the others, producing those Libellas with flat bodies, as Plate 47. Fig. 4. and 6. and Plate 48. Fig. 1. and 5. These are, of all others, the most disagreeable in their appearance; most of them having their bodies and legs very rough and shaggy, and of the exact colour of mud. Others appear of a dirty green, and very unpleasing hue, a colour that in general reigns among them all; for none can boast of any beautiful appearance while in this state, unless the different shapes here exhibited can be called so. Indeed the beauty and symmetry, so apparent in their complete states, make ample amends for the want of it in this; all of them in general being then very beautiful, discovering colours superior to art. [Subfamily, Libellulides. Genus, Libellula.] "The caterpillars of the third sort (see Plate 47. Fig. 3.) are very different from the preceding, being the size of the figure, with the abdomen flat at bottom and rounding at top, longer and slenderer than those at Fig. 2. but not so much as those at Fig. 1. 1. These produce those large Libellas with long slender bodies, who sit at rest with their wings expanded, as Plate 47. Fig. 5. [Subfamily, Libellulides. Genus, Æshna.] "Under these three forms are all the caterpillars of Libellas found, there being but little difference in the colours or marks of the respective tribes; the various sizes and shapes being the chief observable circumstances attending them. They are all furnished with six legs, and have each of them four little membranaceous substances issuing from the back, or upper part of the thorax, that are the follicles, or cases, wherein the wings are inclosed. When the young caterpillars issue from their confinement in the eggs, there is no appearance of these cases, nor till a considerable time after; but as they arrive to a maturer state they become more conspicuous, and, like the young leaves of trees that open "The motion of these creatures in the water, particularly those of the largest size, is very slow; seldom exercising any swiftness or activity, unless they are disturbed and threatened with danger, in which case they can transport themselves to places of more security with the quickness or agility of a fish; but, in general, they appear to have so little inclination to move, that I have often seen them (I mean those that I have kept in glass bowls) remaining in their respective places above a week together, and could not discover the least motion in them, unless under the circumstance above mentioned, or when they had seen their prey and were advancing to seize it. The caterpillars of Fig. 1. 1. and 3. generally fix themselves to some little stick or straw, &c. that they find in the water, and there remain, as I said before, without stirring. Those of Fig. 2. are more frequently seen in motion among the thickest part of the roots and plants that grow there, routing and searching for those small animals inhabiting that part, which are their proper food. This aversion to motion, so apparent in those of No. 1. and 3., appears to me to be the effect of an extraordinary cunning and sagacity; and may be considered as the principal means by which they obtain their prey: for while they continue thus motionless in the water, the small animals, who constitute their proper food, approach them with less fear than they would otherwise do, not suspecting their grand enemy lies upon the watch to seize them the moment they come within his reach; but no sooner has their insensibility of danger brought them within a small distance of those destructive weapons (mentioned before, being placed under their mouths) but that very instant they dart upon them with "I could never observe that these caterpillars ever threw off any exuvia, or skin, in their progress from the egg to the complete animal, as most other insects do that live in the open air; neither could I ever perceive any difference between the caterpillars and chrysalis's of this genus in their outward forms[31]: the same voracious behaviour in seizing and devouring their prey, reigning both in one and the other; but that they internally undergo some material alteration, when passing their respective states, is what I do not entertain the least doubt of; as the organs of respiration during the creature's life in the water, appear to me to be under a necessity of receiving a great alteration, when they are to perform the same office in so different a medium as the air; nor can I suppose this business to be done in so quick and sudden a manner, as the short space of time in which the creature would then be passing from the caterpillar state to the perfect one would permit, without having the intestines prepared, as it were, and fitted by some previous change. However, as this is conjecture only, I shall dwell no longer on this head, my present purpose being to point out and describe their general circumstances and behaviour. "When the caterpillars of the respective species have arrived to their full growth, and nature informs them they are to quit their former element of water, for one wherein they are to appear invested with very different powers, they prepare for this extraordinary change; and, creeping up the sticks, straws, or plants they find for their purpose, whose tops grow out of the water, they entirely quit that element, and, stopping at about six or "Hitherto I have considered these creatures only in their infant or incomplete states; wherein the faculties and powers they are endued with, are entirely different from those of their perfect and complete ones. In the former I have described them swimming and grovelling about in the water, preying upon the lesser kinds of insects, being incapable of subsisting for any length of time out of that element; in the latter we must view them capable of flying in the open air, and conveying themselves from place to place. If they are then confined to the limits of a small pond, they are now capable of roving from tree to tree, and from field to field, darting and skimming along with all the rapidity and seeming joy, that a being sensible of, and exulting in its own powers, can be supposed to do; in short, we "I am strongly inclined to believe, that the greater part, if not all the Libellas, are two years in passing from the egg to the complete animal. But as this is a circumstance I do not remember to have seen in any author, I shall not attempt to establish my opinion as an absolute truth. I shall mention my reasons for this belief, and leave it to every person either to confirm or confute it, as his observations shall hereafter give him an opportunity. "In order to do this, it would be necessary I should mention the times when each species first makes its appearance here in England. But as this would not only take up too much of the reader's time, (there being at least twenty-two of them) but it would likewise be impossible to be understood by any, but the experienced adept, unless I gave figures of them all; I shall therefore content myself with observing, that the Libella figured in Plate 47. Fig. 5. though found in America, about New York, &c. is nearly the same with our English one, differing but a very little from it, and which I have inserted chiefly as an instance to illustrate this subject. Ours is the last species but one, that comes forth in the summer, breeding only once a year, appearing about the 2nd of August. Therefore, to elucidate this point, suppose we allow a fortnight or three weeks from that time for all the Libellas of this species to make their appearance in, that nature intended should be bred that year; but in order to set this matter in the strongest light, I shall allow a month to that purpose. If, therefore, the Libellas were only one year in passing from the egg to the perfect state, we might justly conclude, that after the 2nd of September, (a month from the time of their first appearing) all of this species would have arrived to their complete states; and that none of their caterpillars could be found in the waters after that time, by reason they had all quitted that element, and were become flying insects. This, I say, is the conclusion we might fairly, and without presuming on the matter, make. But this is "These reasons appear to me so convincing, I should have thought myself inexcuseable to have passed them over in silence, when I was giving the history of these animals. I shall therefore think myself happy to have this circumstance corroborated by future experience, or rectified and cleared up, if found to be an error. I shall only say, I have dwelt the longer on it, as it is an observation I never heard of before. I have likewise singled out the largest Libella, and the flat-bodied ones for its illustration, as being more conspicuous by their size, and easier to be observed than those of the smaller sorts. "The two principles of hunger and lust, so apparent through the animal kingdom, are in no class or tribe more manifest and visible than in these insects. The former I have described and explained through their different states; it remains for me to shew the manner in which they obey the calls of the latter. And herein I must observe, that the different manner in which the act of copulation is performed, depends on the difference of the respective kinds; the organs of generation being placed in different parts of the body, according to the distinct species. All the flat-bodied ones have those parts placed in common with most other insects, at the extremity of the tail. In all the slender-bodied ones, the organ of the male is placed next the breast, close to the part where the thorax and abdomen unite; while the same organ in the female lies in the very extremity of the tail, and the singular manner of these creatures coupling is a circumstance worthy of observation. "As soon as they have arrived to their perfect state, the males seek out their mates, in order to propagate their species; for this purpose they frequent ponds and standing waters, places where the females generally harbour, and when the male in the course of his flight comes within a certain distance of her, if it be one of that species whose sexual parts "In this manner all the flat-bodied Libellas copulate and lay their eggs; the others, which are formed with slender bodies, behave very differently in every circumstance; for in each of those species, when the male has singled out its mate, he flies to her, and by means of two little bony substances placed at the end of his tail, issuing on each side, and composing a kind of forceps, he fixes himself to the forepart of her neck, close behind the head, the female discovering no sign of fear while he is performing this action; having fixed himself in this position, he remains there several days, sometimes appearing with his body quite erect, at other times bending himself, and settling with his legs on the same substance she may happen to alight on, without ever quitting his hold; but flies through the air, thus united, wherever the fancy or inclination of the female disposes her to go. This behaviour we must consider as a prelude to copulation, for that is not performed till a considerable time after they are thus united; and therefore I look on it as a wooing, or act of courtship in the male. But having remained a sufficient time in this manner, that is, till the female is disposed to receive him, she bends her tail round to that part where I described the organ of generation to be placed in the male, and, being still held fast by the neck, in that attitude they perform the mandate of nature. This action being over, that in the smaller kinds takes up a considerable space of time (for I have known them in this posture above eighteen hours) the female soon after begins to lay her eggs. She flies to some rush, reed, or other plant growing in the water, and settling close to its edge dips "In this manner all the slender-bodied tribe behave, differing only in some particular circumstances; as, for instance, the largest Libellas, as that at Fig. 5. Pl. 47. are not near so long in copulating or wooing as the small ones; for by the former this act is performed in a very short space, and while they are flying about in the air. Others, of a smaller size, are less quick in this performance; and as we descend to the smallest species, we shall find they take by much the longest time; observing, by the way, that all these slender-bodied ones lay their eggs in the same manner, that is, by fixing them to some substance to which they adhere, till they are ripened into life. My experience well informing me they never scatter them in any loose careless manner as some insects do, but are placed in such proper and apt situations as to receive the influence of the sun to bring them to maturity. "If we attentively consider these creatures, either in their caterpillar or complete states, we cannot help concluding them to be a rank of beings of greater benefit and advantage to mankind than they appear to be at first view; for, not to mention their being annual 'ministers of nature,' they are appointed by the great Governor of the universe as grand instruments for assisting to preserve that equilibrium so apparently reigning through the insect world, and which all who have made any progress in the study of natural history unanimously confess. Hence the voracious disposition of the Libellas is wisely made to answer a most necessary and beneficial purpose; and the great numbers of small insects which are daily sacrificed to their insatiable appetites, both in their caterpillar as well as complete states, is as strong an instance as any I know of the necessity and propriety of the existence of these animals. The general principle reigning through the whole animal kingdom, of the stronger preying upon the weak and defenceless, can hardly be explained to the purpose of being useful to mankind, and agreeable to the laws of nature, in any one instance more than is evidently to be observed in the subjects I have been describing. The least reflection will confirm this; for if the food of the Libellas when in their complete forms had consisted of the leaves of plants, like the locust genus, and not of those small insects they now prey on; or had it, like the beetle tribe, consisted of the superfluous parts of nature, as the putrid carcases of dead animals, rotten wood, &c. how great a chasm would there have been in the universal chain? how evident and conspicuous would it have appeared? and how could the vast number of small insects, increasing every day during the summer, be restrained and lessened? what genus of the transparent-winged class could possibly have performed this business singly? or could all the genera of flies, and even birds that we know of, have accomplished this end? could all the DipterÆ, or any other "Nor is this the only advantage arising from their existence. The still waters, where these creatures are inhabitants during their infant state, are in some degree by their assistance preserved sweet and good, that otherwise might corrupt and putrify, for the motion the waters receive by these insects is not trifling; the respiration they perform in that element being observable by a close attention, which, together with that and the motion of thousands of other insects, does in some measure contribute to keep it sweet and wholesome." |