(8) It appears from the above that mountains and continental elevations may be of three kinds, (a) They may consist of material thrown out of volcanic rents, like earth out of a mole burrow. Mountains like Vesuvius and Ætna are of this kind. (b) They may be parts of wide ridges or chains variously cut and modified by rains and rivers. The Lebanon and the Catskill Mountains are cases in point, (c) They may be lines of crumpling by lateral pressure. The greatest mountains, like the Cordillera, the Alps, and the Appalachians are of this kind, and such mountains may represent lateral pressure occurring at various times, and whose results have been greatly modified subsequently.

I wish to formulate these principles as distinctly as possible, and as the result of all the long series of observations, calculations, and discussions since the time of Werner and Hutton, and in which a vast number of able physicists and naturalists have borne a part, because they may be considered as certain deductions from our actual knowledge, and because they lie at the foundation of a rational physical geology.

We may roughly popularise these deductions by comparing the earth to a drupe or stone-fruit, such as a plum or peach somewhat dried up. It has a large and intensely hard stone and kernel, a thin pulp made up of two layers, an inner, more dense and dark-coloured, and an outer, less dense and lighter-coloured. These constitute the under-crust. On the outside it has a thin membrane or over-crust. In the process of drying it has slightly shrunk, so as to produce ridges and hollows of the outer crust, and this outer crust has cracked in some places, allowing portions of the pulp to ooze out—in some of them its lower dark substance, in others, its upper and lighter material. The analogy extends no farther, for there is nothing in our withered fruit to represent the oceans occupying the lower parts of the surface, or the deposits which they have laid down.

Here a most important feature demands attention. The rain, the streams, and the sea are constantly cutting down the land and depositing it in the bed of the waters. Thus weight is taken from the land, and added to the sea bed. Geological facts, such as the great thickness of the coal measures, in which we find thousands of feet of sediment, all of which must have been deposited in shallow water, and the accumulation of hundreds of feet of superficial material in deltas at the mouth of great rivers, show that the crust of the earth is so mobile as to yield downward to every pressure, however slight.[29] It may do this slowly and gradually, or by jumps from time to time; and this yielding necessarily tends to squeeze up the edges of the depressed portions into ridges, and to cause lateral movement and ejection of volcanic matter at intervals.

Keeping in view these general conclusions, let us now turn to their bearing on the origin and history of the North Atlantic.

Though the Atlantic is a deep ocean, its basin does not constitute so much a depression of the crust of the earth as a flattening of it, and this, as recent soundings have shown, with a slight ridge or elevation along its middle, and banks or terraces fringing the edges, so that its form is not so much that of a basin as that of a shallow elongated plate with its middle a little raised. Its true margins are composed of portions of the over-crust folded, overlapped and crushed, as if by lateral pressure emanating from the sea itself. We cannot, for example, look at a geological map of America without perceiving that the Appalachian ridges, which intervene between the Atlantic and the St. Lawrence valley, have been driven bodily back by a force acting from the east, and that they have resisted this pressure only where, as in the Gulf of St. Lawrence and the Catskill region of New York, they have been protected by outlying masses of very old rocks, as, for example, by that of the island of Newfoundland and that of the Adirondack Mountains. The admirable work begun by my friend and fellow-student, Professor James Nicol, followed up by Professor Lapworth, and now, after long controversy, fully confirmed by the recent observations of the Geological Survey of Scotland, has shown the most intense action of the same kind on the east side of the ocean in the Scottish highlands; and the more widely distributed Eozoic and other old rocks of Scandinavia may be appealed to in further evidence of this.[30]

If we now inquire as to the cause of the Atlantic depression, we must go back to the time when the areas occupied by the Atlantic and its bounding coasts were parts of the shoreless sea in which the earliest gneisses or stratified granites of the Laurentian age were being laid down in vastly extended beds. These ancient crystalline rocks have been the subject of much discussion and controversy, to which reference has been made in a previous chapter.

It will be observed, in regard to these theories, that they do not suppose that the old gneiss is an ordinary sediment, but that all regard it as formed in exceptional circumstances, these circumstances being the absence of land and of subaËrial decay of rock, and the presence wholly or principally of the material of the upper surface of the recently hardened crust. This being granted, the question arises, Ought we not to combine the several theories as to the origin of gneiss, and to believe that the cooling crust has hardened in successive layers from without inward; that at the same time fissures were locally discharging igneous matter to the surface; that matter held in suspension in the ocean and matter held in solution by heated waters rising from beneath the outer crust were mingling their materials in the deposits of the primitive ocean?[31] It would seem that the combination of all these agencies may safely be evoked as causes of the pre-Atlantic deposits. This is the eclectic position I have maintained in a previous chapter, and which I hold to be in every way the most probable.

Let us suppose, then, the floor of old ocean covered with a flat pavement of gneiss, or of that material which is now gneiss, the next question is, How and when did this original bed become converted into sea and land? Here we have some things certain, others most debatable. That the cooling mass, especially if it was sending out volumes of softened rocky material, either in the form of volcanic ejections or in that of matter dissolved in heated water, and piling this on the surface, must soon become too small for its shell, is apparent; but when and where would the collapse, crushing and wrinkling inevitable from this cause begin? The date is indicated by the lines of old mountain chains which traverse the Laurentian districts; but the reason why is less apparent. The more or less unequal cooling, hardening and conductive power of the outer crust we may readily assume. The driftage unequally of water-borne detritus to the south-west by the bottom currents of the sea is another cause, and, as we shall soon see, most effective. Still another is the greater cooling and hardening of the crust in the polar regions, and the tendency to collapse of the equatorial protuberance from the slackening of the earth's rotation. Besides these, the internal tides of the earth's substance at the times of solstice would exert an oblique pulling force on the crust, which might tend to crack it along diagonal lines. From whichever of these causes, or the combination of the whole, we know that, within the Laurentian time, folded portions of the earth's crust began to rise above the general surface, in broad belts running from north-east to south-west, and from north-west to south-east, where the older mountains of Eastern America and Western Europe now stand, and that the subsidence of the oceanic areas, allowed by this crumpling of the crust, permitted other areas on both sides of the Atlantic to form limited table-lands. This was the commencement of a process repeated again and again in subsequent times, and which began in the middle Laurentian, when for the first time we find beds of quartzite, limestone, and iron ore, and graphite beds, indicating that there was already land and water, and that the sea, and perhaps the land, swarmed with forms of animal and plant life, unknown, for the most part, now. Independently of the questions as to the animal nature of Eozoon, I hold that we know, as certainly as we can know anything inferentially, the existence of these primitive forms of life. If I were to conjecture what were these early forms of plant and animal life, still unknown to us by actual specimens, I would suppose that, just as in the PalÆozoic, the acrogens culminated in gigantic and complex forest trees, so in the Laurentian, the algÆ, the lichens, and the mosses grew to dimensions and assumed complexity of structure unexampled in later times, and that, in the sea, the humbler forms of Protozoa and Sea Mosses were the dominant types, but in gigantic and complex forms. The land of this period was probably limited, for the most part, to high latitudes, and its aspect, though more rugged and abrupt, and of greater elevation, must have been of that character which we still see in the Laurentian hills. The distribution of this ancient land is indicated by the long lines of old Laurentian rock extending from the Labrador coast and the north shore of the St. Lawrence, and along the eastern slopes of the Appalachians in America, and the like rocks of the Hebrides, the Western Highlands, and the Scandinavian mountains. A small but interesting remnant is that in the Malvern Hills, so well described by Holl. It will be well to note here, and to fix on our minds, that these ancient ridges of Eastern America and Western Europe have been greatly denuded and wasted since Laurentian times, and that it is along their eastern sides that the greatest sedimentary accumulations have been deposited.

From this time dates the introduction of that dominance of existing causes which forms the basis of uniformitarianism in geology, and which had to go on with various and great modifications of detail, through the successive stages of the geological history, till the land and water of the northern hemisphere attained to their present complex structure.

So soon as we have a circumpolar belt or patches of Eozoic[32] land and ridges running southward from it, we enter on new and more complicated methods of growth of the continents and seas. Portions of the oldest crystalline rocks, raised out of the protecting water, were now eroded by atmospheric agents, and especially by the carbonic acid, then existing in the atmosphere perhaps more abundantly than at present, under whose influence the hardest of the gneissic rocks gradually decay. The arctic lands were subjected, in addition, to the powerful mechanical force of frost and thaw. Thus every shower of rain and every swollen stream would carry into the sea the products of the waste of land, sorting them into fine clays and coarser sands; and the cold currents which cling to the ocean bottom, now determined in their courses, not merely by the earth's rotation, but also by the lines of folding on both sides of the Atlantic, would carry south-westward, and pile up in marginal banks of great thickness the dÉbris produced from the rapid waste of the land already existing in the Arctic regions. The Atlantic, opening widely to the north, and having large rivers pouring into it, was, especially, the ocean characterised, as time advanced, by the prevalence of these phenomena. Thus, throughout the geological history it has happened that, while the middle of the Atlantic has received merely organic deposits of shells of foraminifera and similar organisms, and this probably only to a small amount, its margins have had piled upon them beds of detritus of immense thickness. Professor Hall, of Albany, was the first geologist who pointed out the vast cosmic importance of these deposits, and that the mountains of both sides of the Atlantic owe their origin to these great lines of deposition, along with the fact, afterwards more fully insisted on by Rogers, that the portions of the crust which received these masses of dÉbris became thereby weighted down and softened, and were more liable than other parts to lateral crushing.

Thus, in the later Eozoic and early PalÆozoic times, which succeeded the first foldings of the oldest Laurentian, great ridges were thrown up, along the edges of which were beds of limestone, and on their summits and sides, thick masses of ejected igneous rocks. In the bed of the central Atlantic there are no such accumulations. It must have been a flat, or slightly ridged, plate of the ancient gneiss, hard and resisting, though perhaps with a few cracks, through which igneous matter welled up, as in Iceland and the Azores in more modern times. In this condition of things we have causes tending to perpetuate and extend the distinctions of ocean and continent, mountain and plain, already begun; and of these we may more especially note the continued subsidence of the areas of greatest marine deposition. This has long attracted attention, and affords very convincing evidence of the connection of sedimentary deposit as a cause with the subsidence of the crust.[33]

We are indebted to a French physicist, M. Faye, for an important suggestion on this subject. It is that the sediment accumulated along the shores of the ocean presented an obstacle to radiation, and consequently to cooling of the crust, while the ocean floor, unprotected and unweighted, and constantly bathed with currents of cold water having great power of convection of heat, would be more rapidly cooled, and so would become thicker and stronger. This suggestion is complementary to the theory of Professor Hall, that the areas of greatest deposit on the margins of the ocean are necessarily those of greatest folding and consequent elevation. We have thus a hard, thick, resisting ocean bottom, which, as it settles down toward the interior, under the influence of gravity, squeezes upwards and folds and plicates all the soft sediments deposited on its edges. The Atlantic area is almost an unbroken cake of this kind. The Pacific area has cracked in many places, allowing the interior fluid matter to exude in volcanic ejections.

It may be said that all this supposes a permanent continuance of the ocean basins, whereas many geologists postulate a mid-Atlantic continent to give the thick masses of detritus found in the older formations both in Eastern America and Western Europe, and which thin off in proceeding into the interior of both continents. I prefer, as already stated, to consider these belts of sediment as the deposits of northern currents, and derived from arctic land, and that, like the great banks off the American coast at the present day, which are being built up by the present arctic current, they had little to do with any direct drainage from the adjacent shore. We need not deny, however, that such ridges of land as existed along the Atlantic margins were contributing their quota of river-borne material, just as on a still greater scale the Amazon and Mississippi are doing now, and this especially on the sides toward the present continental plateaus, though the greater part must have been derived from the wide tracts of Laurentian land within the Arctic Circle, or near to it. It is further obvious that the ordinary reasoning respecting the necessity of continental areas in the present ocean basins would actually oblige us to suppose that the whole of the oceans and continents had repeatedly changed places. This consideration opposes enormous physical difficulties to any theory of alternations of the oceanic and continental areas, except locally at their margins.

But the permanence of the Atlantic depression does not exclude the idea of successive submergences of the continental plateaus and marginal slopes, alternating with periods of elevation, when the ocean retreated from the continents and contracted its limits. In this respect the Atlantic of to-day is much smaller than it was in those times when it spread widely over the continental plains and slopes, and much larger than it has been in times of continental elevation. This leads us to the further consideration that, while the ocean beds have been sinking, other areas have been better supported, and constitute the continental plateaus; and that it has been at or near the junctions of these sinking and rising areas that the thickest deposits of detritus, the most extensive foldings, and the greatest ejections of volcanic matter have occurred. There has thus been a permanence of the position of the continents and oceans throughout geological time, but with many oscillations of these areas, producing submergences and emergences of the land. In this way we can reconcile the vast vicissitudes of the continental areas in different geological periods with that continuity of development from north to south, and from the interiors to the margins, which is so marked a feature. We have, for this reason, to formulate another apparent geological paradox, namely, that while, in one sense, the continental and oceanic areas are permanent, in another, they have been in continual movement. Nor does this view exclude extension of the continental borders or of chains of islands beyond their present limits, at certain periods; and indeed, the general principle already stated, that subsidence of the ocean bed has produced elevation of the land, implies in earlier periods a shallower ocean and many possibilities as to volcanic islands, and low continental margins creeping out into the sea; while it is also to be noted that there are, as already stated, bordering shelves, constituting shallows in the ocean, which at certain periods have emerged as land.

We are thus compelled, as already stated, to believe in the contemporaneous existence in all geological periods, except perhaps the earliest of them, of the three distinct conditions of areas on the surface of the earth, defined in chapter second oceanic areas of deep sea, continental plateaus and marginal shelves, and lines of plication and folding.

In the successive geological periods the continental plateaus, when submerged, owing to their vast extent of warm and shallow sea, have been the great theatres of the development of marine life and of the deposition of organic limestones, and when elevated, they have furnished the abodes of the noblest land faunas and floras. The mountain belts, especially in the north, have been the refuge and stronghold of land life in periods of submergence; and the deep ocean basins have been the perennial abodes of pelagic and abyssal creatures and the refuge of multitudes of other marine animals and plants in times of continental elevation. These general facts are full of importance with reference to the question of the succession of formations and of life in the geological history of the earth.

So much space has been occupied with these general views, that it would be impossible to trace the history of the Atlantic in detail through the ages of the PalÆozoic, Mesozoic, and Tertiary. We may, however, shortly glance at the changes of the three kinds of surface already referred to. The bed of the ocean seems to have remained, on the whole, abyssal; but there were probably periods when those shallow reaches of the Atlantic which stretch across its most northern portion, and partly separate it from the Arctic basin, presented connecting coasts or continuous chains of islands sufficient to permit animals and plants to pass over.[34] At certain periods also there were, not unlikely, groups of volcanic islands, like the Azores, in the temperate or tropical Atlantic. More especially might this be the case in that early time when it was more like the present Pacific; and the line of the great volcanic belt of the Mediterranean, the mid-Atlantic banks, the Azores and the West India Islands point to the possibility of such partial connections. These were stepping stones, so to speak, over which land organisms might cross, and some of these may be connected with the fabulous or pre-historic Atlantis.

In the PalÆozoic period, the distinctions already referred to, into continental plateaus, mountain ridges, and ocean depths, were first developed, and we find, already, great masses of sediment accumulating on the seaward sides of the old Laurentian ridges, and internal deposits thinning away from these ridges over the submerged continental areas, and presenting dissimilar conditions of sedimentation. It would seem also that, as Hicks has argued for Europe, and Logan and Hall for America, this Cambrian age was one of slow subsidence of the land previously elevated, accompanied with or caused by thick deposits of detritus along the borders of the subsiding shore, which was probably covered with the decomposing rock arising from long ages of subaËrial waste.

In the coal formation age its characteristic swampy flats stretched in some places far into the shallower parts of the ocean.[35] In the Permian, the great plicated mountain margins were fully developed on both sides of the Atlantic. In the Jurassic, the American continent probably extended farther to the sea than at present. In the Wealden age there was much land to the west and north of Great Britain, and Professor Bonney has directed attention to the evidence of the existence of this land as far back as the Trias, while Mr. Starkie Gardiner has insisted on connecting links to the southward, as evidenced by fossil plants. So late as the Post-glacial, or early human period, large tracts, now submerged, formed portions of the continents. On the other hand, the interior plains of America and Europe were often submerged. Such submergences are indicated by the great limestones of the PalÆozoic, by the chalk and its representative beds in the Cretaceous, by the Nummulitic formation in the Eocene, and lastly, by the great Pleistocene submergence, one of the most remarkable of all, one in which nearly the whole northern hemisphere participated, and which was probably separated from the present time by only a few thousands of years.[36] These submergences and elevations were not always alike on the two sides of the Atlantic. The Salina period of the Silurian, for example, and the Jurassic, show continental elevation in America not shared by Europe. The great subsidences of the Cretaceous and the Eocene were proportionally deeper and wider on the eastern continent, and this and the direction of the land being from north to south, cause more ancient forms of life to survive in America. These elevations and submergences of the plateaus alternated with the periods of mountain-making plication, which was going on at intervals, at the close of the Eozoic, at the beginning of the Cambrian, at the close of the Siluro-Cambrian, in the Permian, and in Europe and Western America in the Tertiary. The series of changes, however, affecting all these areas was of a highly complex character in detail.[37]

We may also note a fact which I have long ago insisted on,[38] the regular pulsation of the continental areas, giving us alternations in each great system of deep-sea and shallow-water beds, so that the successive groups of formations may be divided into triplets of shallow-water, deep-water, and shallow-water strata, alternating in each period. This law of succession applies more particularly to the formations of the continental plateaus, rather than to those of the ocean margins, and it shows that, intervening between the great movements of plication there were subsidences of those plateaus, or elevations of the sea bottom, which allowed the waters to spread themselves over all the inland spaces between the great folded mountain ranges of the Atlantic borders.

In referring to the ocean basins we should bear in mind that there are three of these in the northern hemisphere the Arctic, the Pacific, and the Atlantic. De Ranee has ably summed up the known facts as to Arctic geology in a series of articles in Nature, from which it appears that this area presents from without inwards a succession of older and newer formations from the Eozoic to the Tertiary, and that its extent must have been greater in former periods than at present, while it must have enjoyed a comparatively warm climate from the Cambrian to the Pleistocene period. The relations of its deposits and fossils are closer with those of the Atlantic than with those of the Pacific, as might be anticipated from its wider opening into the former. Blandford has recently remarked on the correspondence of the marginal deposits around the Pacific and Indian oceans,[39] and Dr. Dawson informs me that this is equally marked in comparison with the west coast of America, but these marginal areas have not yet gained much on the ocean. In the North Atlantic, on the other hand, there is a wide belt of comparatively modern rocks on both sides, more especially toward the south and on the American side; but while there appears to be a perfect correspondence on both sides of the Atlantic, and around the Pacific respectively, there seems to be less parallelism between the deposits and forms of life of the two oceans, as compared with each other, and less correspondence in forms of life, especially in modern times. Still, in the earlier geological ages, as might have been anticipated from the imperfect development of the continents, the same forms of life characterise the whole ocean from Australia to Arctic America, and indicate a grand unity of Pacific and Atlantic life not equalled in later times,[40] and which speaks of true contemporaneity rather than of what has been termed homotaxis or mere likeness of orders.

We may pause here for a moment to notice some of the effects of Atlantic growth on modern geography. It has given us rugged and broken shores, composed of old rocks in the north, and newer formations and softer features toward the south. It has given us marginal mountain ridges and internal plateaus on both sides of the sea. It has produced certain curious and by no means accidental correspondences of the eastern and western sides. Thus the solid basis on which the British Islands stand may be compared with Newfoundland and Labrador, the English Channel with the Gulf of St. Lawrence, the Bay of Biscay with the Bay of Maine, Spain with the projection of the American land at Cape Hatteras, the Mediterranean with the Gulf of Mexico. The special conditions of deposition and plication necessary to these results, and their bearing on the character and productions of the Atlantic basin, would require a volume for their detailed elucidation.

Thus far our discussion has been limited almost entirely to physical causes and effects. If we now turn to the life history of the Atlantic, we are met at the threshold with the question of climate, not as a thing fixed and immutable, but as changing from age to age in harmony with geographical mutations, and producing long cosmic summers and winters of alternate warmth and refrigeration.

We can scarcely doubt that the close connection of the Atlantic and Arctic oceans is one factor in those remarkable vicissitudes of climate experienced by the former, and in which the Pacific area has also shared in connection with the Antarctic Sea. No geological facts are indeed at first sight more strange and inexplicable than the changes of climate in the Atlantic area, even in comparatively modern periods. We know that in the early Tertiary temperate conditions reigned as far north as the middle of Greenland, and that in the Pleistocene the Arctic cold advanced until an almost perennial winter prevailed half way to the equator. It is no wonder that nearly every cause available in the heavens and the earth has been invoked to account for these astounding facts. I shall, I trust, be excused if, neglecting most of these theoretical views, I venture to invite attention, in connection with this question, chiefly to the old Lyellian doctrine of the modification of climate by geographical changes. Let us, at least, consider how much these are able to account for.

The ocean is a great equalizer of extremes of temperature. It does this by its great capacity for heat, and by its cooling and heating power when passing from the solid into the liquid and gaseous states, and the reverse. It also acts by its mobility, its currents serving to convey heat to great distances, or to cool the air by the movement of cold icy waters. The land, on the other hand, cools or warms rapidly, and can transmit its influence to a distance only by the winds, and the influence so transmitted is rather in the nature of a disturbing than of an equalizing cause. It follows that any change in the distribution of land and water must affect climate, more especially if it changes the character or course of the ocean currents.

Turning to the Atlantic, in this connection we perceive that its present condition is peculiar and exceptional. On the one hand it is widely open to the Arctic Sea and the influence of its cold currents, and on the other it is supplied with a heating apparatus of enormous power to give a special elevation of temperature, more particularly to its eastern coasts. The great equatorial current running across from Africa is on its northern side embayed in the Gulf of Mexico, as in a great cauldron, and pouring through the mouth of this in the Bahama channel, forms the gulf stream, which, widening out like a fan, forms a vast expanse of warm water, from which the prevailing westerly winds of the North Atlantic waft a constant supply of heated moist air to the western coasts of Europe, giving them a much more warm and uniform climate than that which prevails in similar latitudes in Eastern America, where the cold Arctic currents hug the shore, and bring down ice from Baffin's Bay. Now all this might be differently arranged. We shall find that there were times, when the Isthmus of Panama being broken through, there was no Gulf Stream, and Norway and England were reduced to the conditions of Greenland and Labrador, and when refrigeration was still further increased by subsidence of northern lands affording freer sweep to the Arctic currents. On the other hand, there were times when the Gulf of Mexico extended much farther north than at present, and formed an additional surface of warm water to heat all the interior of America, as well as the Atlantic. Geographical changes of these kinds, have probably given us the glacial period in very recent times, and at an earlier era those warm climates which permitted temperate vegetation to flourish as far north as Greenland. These are, however, great topics, which must form the subject of other chapters.

I am old enough to remember the sensation caused by the delightful revelations of Edward Forbes respecting the zones of animal life in the sea, and the vast insight which they gave into the significance of the work on minute organisms previously done by Ehrenberg, Lonsdale and Williamson, and into the meaning of fossil remains. A little later the soundings for the Atlantic cable revealed the chalky foraminiferal ooze of the abyssal ocean. Still more recently, the wealth of facts disclosed by the Challenger voyage, which naturalists have scarcely yet had time to digest, have opened up to us new worlds of deep-sea life.

The bed of the deep Atlantic is covered, for the most part, by a mud or ooze, largely made up of the dÉbris of foraminifera and other minute organisms mixed with fine clay. In the North Atlantic the Norwegian naturalists call this the Biloculina mud. Farther south, the Challenger naturalists speak of it as Globigerina ooze. In point of fact it contains different species of foraminiferal shells, Globigerina and Orbulina being in some localities dominant, and in others, other species; and these changes are more apparent in the shallower portions of the ocean.

On the other hand, there are means for disseminating coarse material over parts of the ocean beds. There are, in the line of the Arctic current, on the American coast, great sand banks, and off the coast of Norway, sand constitutes a considerable part of the bottom material. Soundings and dredgings off Great Britain, and also off the American coast, have shown that fragments of stone referable to Arctic lands are abundantly strewn over the bottom, along certain lines, and the Antarctic continent, otherwise almost unknown, makes its presence felt to the dredge by the abundant masses of crystalline rock drifted far from it to the north. These are not altogether new discoveries. I had inferred, many years ago, from stones taken up by the hooks of fishermen on the banks of Newfoundland, that rocky material from the north is dropped on these banks by the heavy ice which drifts over them every spring, that these are glaciated, and that after they fall to the bottom sand is drifted over them with sufficient velocity to polish the stones, and to erode the shelly coverings of Arctic animals attached to them.[41] If, then, the Atlantic basin were upheaved into land, we should see beds of sand, gravel and boulders with clay flats and layers of marl and limestone. According to the Challenger reports, in the Antarctic seas S. of 64 there is blue mud, with fragments of rock, in depths of 1,200 to 2,000 fathoms. The stones, some of them glaciated, were granite, diorite, amphibolite, mica schist, gneiss and quartzite. This deposit ceases and gives place to Globigerina ooze and red clay at 46° to 47° S., but even farther north there is sometimes as much as 49 per cent, of crystalline sand. In the Labrador current a block of syenite, weighing 400 lbs., was taken up from 1,340 fathoms, and in the Arctic current, 100 miles from land, was a stony deposit, some stones being glaciated. Among these were smoky quartz, quartzite, limestone, dolomite, mica schist, and serpentine; also particles of monoclinic and triclinic felspar, hornblende, augite, magnetite, mica and glauconite, the latter, no doubt, formed in the sea bottom, the others drifted from Eozoic and PalÆozoic formations to the north.[42]

A remarkable fact in this connection is that the great depths of the sea are as impassable to the majority of marine animals as the land itself. According to Murray, while twelve of the Challenger's dredgings, taken in depths greater than 2,000 fathoms, gave 92 species, mostly new to science, a similar number of dredgings in shallower water near the land, give no less than 1,000 species. Hence arises another apparent paradox relating to the distribution of organic beings. While at first sight it might seem that the chances of wide distribution are exceptionally great for marine species, this is not so. Except in the case of those which enjoy a period of free locomotion when young, or are floating and pelagic, the deep ocean sets bounds to their migrations. On the other hand, the spores of cryptogamic plants may be carried for vast distances by the wind, and the growth of volcanic islands may effect connections which, though only temporary, may afford opportunity for land animals and plants to pass over.

With reference to the transmission of living beings across the Atlantic, we have before us the remarkable fact that from the Cambrian age onwards there were, on the two sides of the ocean, many species of invertebrate animals which were either identical or so closely allied as to be possibly varietal forms, indicating probably the shallowness of the ocean in these periods. In like manner, the early plants of the Upper Silurian, Devonian, and Carboniferous present many identical species; but this identity is less marked in more modern times. Even in the latter, however, there are remarkable connections between the floras of oceanic islands and the continents. Thus the Bermudas, altogether recent islands, have been stocked by the agency chiefly of the ocean currents and of birds, with nearly 150 species of continental plants; and the facts collected by Helmsley as to the present facilities of transmission, along with the evidence afforded by older oceanic islands which have been receiving animal and vegetable colonists for longer periods, go far to show that, time being given, the sea actually affords facilities for the migration of the inhabitants of the land, comparable with those of continuous continents.

In so far as plants are concerned, it is to be observed that the early forests were largely composed of cryptogamous plants, and the spores of these in modern times have proved capable of transmission from great distances. In considering this, we cannot fail to conclude, that the union of simple cryptogamous fructification with arboreal stems of high complexity, so well illustrated by Dr. Williamson, had a direct relation to the necessity for a rapid and wide distribution of these ancient trees. It seems also certain that some spores, as, for example, those of the Rhizocarps,[43] a type of vegetation abundant in the PalÆozoic, and certain kinds of seeds, as those named Æthoetesta and Pachytheca, were fitted for flotation. Further, the periods of Arctic warmth permitted the passage around the northern belt of many temperate species of plants, just as now happens with the Arctic flora; and when these were displaced by colder periods, they marched southward along both sides of the sea on the mountain chains.

The same remark applies to northern forms of marine invertebrates, which are much more widely distributed in longitude than those farther south. The late Mr. Gywn Jeffreys, in one of his latest communications on this subject, stated that 54 per cent, of the shallow-water mollusks of New England and Canada are also European, and of the deep-sea forms, 30 out of 35; these last, of course, enjoying greater facilities for migration than those which have to travel slowly along the shallows of the coast in order to cross the ocean and settle themselves on both sides. Many of these animals, like the common mussel and sand clam, are old settlers which came over in the Pleistocene period, or even earlier. Others, like the common periwinkle, seem to have been slowly extending themselves in modern times, perhaps even by the agency of man. The older immigrants may possibly have taken advantage of lines of coast now submerged, or of warm periods, when they could creep round the Arctic shores. Mr. Herbert Carpenter and other naturalists employed on the Challenger collections have made similar statements respecting other marine invertebrates, as, for instance, the Echinoderms, of which the deep-sea crinoids present many common species, and my own collections prove that many of the shallow-water forms are common. Dall and Whiteaves[44] have shown that some mollusks and Echinoderms are common even to the Atlantic and Pacific coasts of North America; a remarkable fact, testifying at once to the fixity of these species and to the manner in which they have been able to take advantage of geographical changes. Some of the species of whelks common to the Gulf of St. Lawrence and the Pacific are animals which have no special locomotive powers, even when young, but they are northern forms not proceeding far south, so that they may have passed through the Arctic seas. In this connection it is well to remark that many species of animals have powers of locomotion in youth which they lose when adult, and that others may have special means of transit. I once found at GaspÉ a specimen of the Pacific species of Coronula, or whale-barnacle, the C. reginÆ of Darwin, attached to a whale taken in the Gulf of St. Lawrence, and which had possibly succeeded in making that passage around the north of America which so many navigators have essayed in vain.[45]

[44] Dall, Report on Alaska; Whiteaves, Trans. R. S. C.