Love out their hour and leave their lives in air. CANTO IV. l. 456. From the accurate experiments and observations of Spallanzani it appears that in the Spartium Junceum, rush-broom, the very minute seeds were discerned in the pod at least twenty days before the flower is in full bloom, that is twenty days before fecundation. At this time also the powder of the anthers was visible, but glued fast to their summits. The seeds however at this time, and for ten days after the blossom had fallen off, appeared to consist of a gelatinous substance. On the eleventh day after the falling of the blossom the seeds became heart- shape, with the basis attached by an appendage to the pod, and a white point at the apex; this white point was on pressure found to be a cavity including a drop of liquor. On the 25th day the cavity which at first appeared at the apex was much enlarged and still full of liquor, it also contained a very small semi- transparent body, of a yellowish colour, gelatinous, and fixed by its two opposite ends to the sides of the cavity. In a month the seed was much enlarged and its shape changed from a heart to a kidney, the little body contained in the cavity was increased in bulk and was less transparent, and gelatinous, but there yet appeared no organization. On the 40th day the cavity now grown larger was quite filled with the body, which was covered with a thin membrane; after this membrane was removed the body appeared of a bright green, and was easily divided by the point of a needle into two portions, which manifestly formed the two lobes, and within these attached to the lower part the exceedingly small plantule was easily perceived. The foregoing observations evince, 1. That the seeds exist in the ovarium many days before fecundation. 2. That they remain for some time solid, and then a cavity containing a liquid is formed in them. 3. That after fecundation a body begins to appear within the cavity fixed by two points to the sides, which in process of time proves to be two lobes containing a plantule. 4. That the ripe seed consists of two lobes adhering to a plantule, and surrounded by a thin membrane which is itself covered with a husk or cuticle. Spalanzani's Dissertations, Vol. II. p. 253. The analogy between seeds and eggs has long been observed, and is confirmed by the mode of their production. The egg is known to be formed within the hen long before its impregnation; C.F. Wolf asserts that the yolk of the egg is nourished by the vessels of the mother, and that it has from those its arterial and venous branches, but that after impregnation these vessels gradually become impervious and obliterated, and that new ones are produced from the fetus and dispersed into the yolk. Haller's Physiolog. Tom. VIII. p. 94. The young seed after fecundation, I suppose, is nourished in a similar manner from the gelatinous liquor, which is previously deposited for that purpose; the uterus of the plant producing or secreting it into a reservoir or amnios in which the embryon is lodged, and that the young embryon is furnished with vessels to absorb a part of it, as in the very early embryon in the animal uterus. The spawn of frogs and of fish is delivered from the female before its impregnation. M. Bonnet says that the male salamander darts his semen into the water, where it forms a little whitish cloud which is afterwards received by the swoln anus of the female, and she is fecundated.—He adds that marine plants approach near to these animals, as the male does not project a fine powder but a liquor which in like manner forms a little cloud in the water.—And further adds, who knows but the powder of the stamina of certain plants may not make some impression on certain germs belonging to the animal kingdom! Letter XLIII. to Spalanzani, Oevres Philos. Spalanzani found that the seminal fluid of frogs and dogs even when diluted with much water retained its prolific quality. Whether this quality be simply a stimulus exciting the egg into animal action, which may be called a vivifying principle, or whether part of it be actually conjoined with the egg is not yet determined, though the latter seems more probable from the frequent resemblance of the fetus to the male parent. A conjunction however of both the male and female influence seems necessary for the purpose of reproduction throughout all organized nature, as well in hermaphrodite insects, microscopic animals, and polypi, and exists as well in the formation of the buds of vegetables as in the production of their seeds, which is ingeniously conceived and explained by Linneus. After having compared the flower to the larva of a butterfly, confining of petals instead of wings, calyxes instead of wing-sheaths, with the organs of reproduction, and having shewn the use of the farina in fecundating the egg or seed, he proceeds to explain the production of the bud. The calyx of a flower, he says, is an expansion of the outer bark, the petals proceed from the inner bark or rind, the stamens from the alburnum or woody circle, and the style from the pith. In the production and impregnation of the seed a commixture of the secretions of the stamens and style are necessary; and for the production of a bud he thinks the medulla or pith bursts its integuments and mixes with the woody part or alburnum, and these forcing their passage through the rind and bark constitute the bud or viviparous progeny of the vegetable. System of Vegetables translated from Linneus, p. 8. It has been supposed that the embryon vegetable after fecundation, by its living activity or stimulus exerted on the vessels of the parent plant, may produce the fruit or seed-lobes, as the animal fetus produces its placenta, and as vegetable buds may be supposed to produce their umbilical vessels or roots down the bark of the tree. This in respect to the production of the fruit surrounding the seeds of trees has been assimilated to the gall-nuts on oak-leaves, and to the bedeguar on briars, but there is a powerful objection to this doctrine, viz. that the fruit of figs, all which are female in this country, grow nearly as large without fecundation, and therefore the embryon has in them no self-living principle. |