[a] It was at one time supposed that the concretions in the marginal bodies of medusÆ represented lenses and the surrounding nerve tissue the optic nerve, a supposition so highly improbable that it never gained any acceptance. (Ib., p. 41, note.)
[b] Eimer’s results I get from Romanes and Hesse[III].
[c] By no means do I wish to attribute intelligence to these animals.
[d] Haake[2] says that in the adult Charybdea Rostonii the vitreous bodies of the complex eyes are absent but present in the young. It is difficult to explain this observation except on grounds of imperfect preservation of the adult material, for in all observations on other forms a vitreous body is described. Haake evidently did not use sections, and for this reason his results must be regarded as of doubtful accuracy. Haake also says that the simple lateral eyes of the clubs are absent in the adult, but present in the young.
[e] In the series from which Fig. 3 is taken the pyramid-cells are not so readily demonstrated. Indeed, I missed them altogether at first in this and some other series and supposed that there were only two kinds of cells (19), but upon a careful re-examination I could demonstrate them to my satisfaction. They did not show, however, in the particular section of Fig. 3, so that they are not indicated in this figure.
[f] I go into this at some length because the cell-walls in the series that showed the nuclei best differentiated as lighter and darker ones did not show well, and there might be some doubt that these lighter nuclei belonged to the pyramid cells. I could, however, in many instances, trace the axial fibers of the pyramids through the pigmented zone to these lighter nuclei (as already noted) which fact can leave no doubt but that some of these nuclei belong to the pyramid cells. (Similar nuclei, however, are found to belong to the long pigment cells, to be described below.) Centrad these pyramid cells are continued into a single process just as the prism cells were shown to be (Fig. 7). Figures 6, 8, 9, and 21 show samples of all the pigmented cells found in macerated preparations, and none of these (except Fig. 9, long pigment cells) show more than a single centrad process. Hence, I conclude that centrad both the pyramid cells and prism cells are continued as a single prolongation.
[g] I have been able to demonstrate nucleoli in all the different nuclei of the cells of the sensory clubs.
[h] It may be objected that my criterion, the presence of axial fibers, is not necessarily characteristic of visual cells. However, the great general occurrence of such axial fibers (Patten,[5] Grenacher,[16] Schreiner,[12] Hesse,[13] myself, in simple complex eye, see below, and perhaps others) in eyes in which the retina has only one kind of cells, would seem to indicate that they are quite characteristic of visual cells. Note again that in the proximal eye of Charybdea there is only one kind of cells and with axial fibers.
[i] Mr. J. C. Olsen, of the Chemical Laboratory, kindly made these tests for me.
