George Adams

The subject of the following chapter opens an extensive field for observation to the naturalist, in which the labour of a life may be well employed: it is a branch where the observer will find the microscope of continual use, and without which he will scarce be able to form any just idea of the organization of trees and plants, or of the variations in the disposition, the number, nature, and offices of their several parts.

Vegetables are beautiful and perfect in their kind, wonderful in their growth, beneficial in their uses. “Herbs and flowers may be regarded by some persons as objects of inferior consideration in philosophy; but every thing must be great which has God for its author. To him all the parts of nature are equally related: the flowers of the earth can raise our thoughts up to the Creator of the world as effectually as the stars of heaven; and, till we make this use of both, we cannot be said to think properly of either. All trees and herbs in their place and seasons speak the same language from the climates of the north to the torrid regions of the south, and from the winter to the spring and the harvest, they join their voices in the universal chorus of all created beings, and to the ear of reason celebrate the wisdom of the Almighty Creator.”

Malpighi, Grew, Duhamel, Hill, Bonnet, and De Saussure, are almost the only writers who have treated on the interior structure of vegetables; and, if we consider the imperfection of the instruments used by some of them in these anatomical researches, and the little attention paid by the rest to the advantages their favourite pursuits might have derived from the use of the microscope, as well as the dissecting knife, we find greater cause to wonder at what has been done, than at what remains to be performed. To the general inattention to the structure of plants, we may, amongst other causes, also ascribe the instability and fluctuation of the different theories on the principles of vegetation. We are, however, so little acquainted with the steps which Providence takes to lead intellectual, but free agents, to the knowledge of truth, and the various difficulties, errors, and prejudices, necessary to be removed, before it can shine in its native colours, that it is our duty to encourage every humble effort towards the advancement of science, that thus we may co-operate with our Creator and Redeemer in promoting that vast plan to which all things are now converging, the bringing all his creatures to a state of truth, goodness, and consequent happiness, an end worthy of the best and wisest of beings.[129]

[129] See the Bishop of Exeter’s Sermon before the Society for the Propagation of the Gospel.

As Dr. Hill is the first writer who has treated this part of natural history in an orderly and scientific manner, I shall use the names he has adopted for characterizing the different parts of trees, &c. which are, 1. the rind; 2. the bark; 3. the blea; 4. the wood; 5. the corona or circle of propagation; 6. the pith. These are placed immediately within or under one another; they are the essential parts upon which the strength of the tree depends: in, among, and between these, the various vessels are placed, which nourish the whole, and maintain and carry on the vegetation of the tree, and from which it obtains its peculiar qualities and virtues. These vessels are of five kinds:

1.	The Exterior		-	Juice Vessels.
2.	The Interior
3.	The Intimate
4.	The Sap Vessels.
5.	The Coronal.

Of these, the first are placed between the rind and bark; the second, in the substance of the bark; the third, in the substance of the blea; the fourth, in the substance of the wood; the fifth, in the corona.

More accurate instruments, or a more minute investigation of the parts, may probably discover new vessels in a system which appears to be entirely vascular, and brings us more thoroughly acquainted with the nature of vegetation.

OF THE RIND.

The exterior covering of all trees is a thin, dry, parched substance, which has been compared by many writers to the skin of animals, and called by names analogous thereto; thus it is called the epidermis by Duhamel, the skin by Grew, the rind by Hill.

When a tree is full of sap, this membrane may be easily detached from the part it covers; it may be separated from green branches which are not in sap, by boiling them in water; large pieces of it may also be obtained from rotten branches; the rind of the leaves of many trees is detached with singular dexterity from the other parts, by some of the mining caterpillars; artificial methods for effecting this purpose have been described in page 160 of this work. Though the rind may at first sight be thought to be of little use, it will be found to be a principal organ in the process of vegetation. The part which covers the root has the most important offices assigned to it.

Many are of opinion that the rind is formed of dried vesiculÆ; and Malpighi says, that we may see in the vascular texture of the bark of the cherry and plumb-trees an arrangement of the parts proper to form the rind, and this arrangement is occasioned by the endeavour of the vascular part to extend itself to the circumference, and the resistance it meets with from the rind; and that hence the vessels are flattened, and assume a membranaceous form.

The rind is a general covering to the young trunks of trees, to the branches, the roots, the leaves, the fruit, the flowers, &c. Upon the trunks of large trees some pieces only of the rind are to be found, having probably been broken by the increased size of the tree. The rind of some species of trees will bear being stretched much further than those of others, and remain for a considerable time uniformly spread over the bark. Du Hamel asserts, that the rind of vigorous healthy, trees remains longer whole than on those that are more languid, notwithstanding that the growth of the last is slower, and therefore makes less efforts against the rind. This circumstance is much in favour of the distinct organization of the rind, and against the opinion of those who only suppose it to consist of dried bladders.

Thin as the rind is, it is formed of many coats, adhering closely to each other, which in some species may be separated with ease, in others, with difficulty. Du Hamel says, that he has divided the rind of the birch into six distinct coats, and that he had no doubt but what the division might have been carried much further. Dr. Hill says, that unless some of these coats be obtained in a state of separation from the rest, the true construction of the rind cannot be discovered, for the connection and form of the parts are lost by the confusion in which they appear while they lie one upon another.

The following experiments may throw a little light upon this obscure subject.[130] All the rind was taken from the trunk of a cherry-tree, and the tree thus skinned exposed to the air; a part of the bark which was next to the rind dried up and exfoliated; the part next to this did the same; after two or three exfoliations, a farinaceous substance covered the superfice of the trunk, soon after which a new rind appeared. Some pieces of rind were taken from a few young branches, and the wounds were covered with a cloth that had been soaked in wax and turpentine; on these the rind appeared in a very little time, without any apparent exfoliation. From some other branches, not only the rind, but a part of the bark was also taken away, and the wounds covered as before; a slight exfoliation was observed here, which was soon followed with a new rind. The bark was taken entirely off from a vigorous cherry-tree, while it was in full sap, so that the wood appeared the whole extent of the trunk. This was protected from the rays of the sun, and from the air. A new bark and rind formed themselves upon the trunk, but they did not originate from the bark that was left on the branches and the root, but extended from different spots, which were first formed at considerable distances from each other. After a lapse of fifteen years, this new rind did not appear like the natural rind of the cherry-tree. From these experiments we learn, that the rind regenerates more readily in some cases than in others, and that it preserves and prevents in a degree the bark from becoming dry too soon, and in consequence thereof exfoliating.

[130] Du Hamel Physique des Arbres, tom. 1, p. 12.

Aided by the microscope, a number of luminous points may be discovered in the rind;[131] these are so many minute holes for other purposes of transpiration. In the cane these holes are visible to the naked eye. A few oval holes may also be perceived in it; these are, however, no more than a separation of the parts, occasioned by the extension of the vasa interiora.

Dr. Grew supposed the rind to be formed of small vesicles, or bladders, clustered together, and intermixed with ligneous fibres or vessels, which run through the length of the rind; these are conjoined by other transverse ones, but that as the rind dries, the bladders or blebs shrink up and disappear. This account does not differ much from that of Dr. Hill, who says, that the rind is formed of a series of longitudinal vessels, and a filmy substance, between them, which, when viewed in a transverse section, form small circles, the sides of which are supported and made up of these longitudinal fibres; that the transverse vessels are only a deception, occasioned by the spaces between them and part of the film. The mode of obtaining an accurate view of the organization of this part, by conveying coloured liquors into the several vessels thereof, has been already described in page 160 of these Essays; by these means, together with the microscope, we find that the vessels are everywhere pierced with small dots or openings: of the use of these, the following conjectures have been formed.[132]

The root, which is equal in surface to a third part of the tree above ground, is covered with a pierced rind. The cold of winter contracts the whole of this, the parts are drawn closer together, and the mouths of these innumerable vessels are shut or nearly so, by this contraction; a very little of the half-congealed moisture of the ground gets into them, but this suffices for the service of the tree, when there is little heat to cause any perspiration, and at a time when in the deciduous trees, the very organs of the greatest perspiration, the leaves, do not exist.

The warmth of the spring arrives, the fluids of the earth grow thinner, every part of the root expands; this opens the mouths of the vessels, and the torrent of nutrition rushes in. By these means, every coat of the rind, and the interstitial spaces thereof, are rendered supple, and may be easily separated from the under coverings.

In roots, the colour of the rind varies very much, being white in some, brown in others, &c. Every root, according to Grew, after it has arrived at a certain age, has a double skin, the one coeval with the other parts, and exists in the seed; a ring is afterwards sent off from the bark, which forms the second skin; thus in the root of dandelion, towards the end of May, the original or outer skin appears shrivelled, and is easily separated from the new one, which is fresher, and adheres more firmly to the bark. Perennial plants are supplied in this manner with a new skin every year; the outer one always falls off in the autumn and winter, and a new one is formed from the bark in the succeeding spring.

OF THE VESSELS WHICH ARE CONTAINED BETWEEN THE RIND AND THE BARK.

These are called by Du Hamel the cellular coat, enveloppe cellulaire; by Hill, the exterior vessels, and the vasa propria exteriora.

It has been already observed, that in trees the juice vessels, or vasa propria, do not form those constituent parts of the wood of which the timber consists, but that it is from the nature of these recipient vessels that it derives its virtues, qualities, and specific properties.[133] A tree may grow, live, and give shade without them; but on those its peculiar character and decided virtues depend; these are greatest where the vasa propria are largest or most numerous; and where we do not find these, we scarce find any thing that will affect the taste or the smell. There are different ranges of these vessels between the several parts, each of which has its allotted place, its peculiar form, its different structure, and its separate use. Many trees have them in all their parts, others only in some of them, while others do not exhibit any.

On taking off the rind, we find a substance of a deep green colour, succulent and herbaceous, formed of a prodigious number of filaments interwoven together in various directions; it is more abundant in some trees than in others, particularly in the elder, and more succulent in summer than in winter; it is then also less adherent to the rind. Dr. Hill thinks the best time of separating the rind, in order to view this part, is in a living branch, at the time of its swelling for the spring, or for the midsummer shoot, but much easier by the means of maceration.[134]

When the rind is perfectly separated, it leaves the vasa propria of this class behind it; they scarce adhere to the inner bark, and but little to the rind; they are disposed in packets, and do not run straight down the branch, but interweaving with each other, form a kind of net. These packets may be separated easily from the bark; when a thin transverse section of one of them is examined, it is found to be composed of twelve or fifteen distinct vessels with hard rinds. Dr. Hill says, that with a great deal of patience, a vast number of objects, and a good microscope, we may see by what means these vessels adhere to the bark; for we shall find upon the sides small oval depressions which fit thereto, and that are probably a kind of glands, that separate from the general store of sap, with which the bark is filled, the juices peculiar to these vessels.

OF THE BARK.

The bark lies next within the rind, and differs but little from it in construction, though it holds a more important office in the scale of vegetation, the growth and qualities of the tree being in a great measure connected with it. It is, therefore, found to differ considerably in substance, quantity, and quality, in various kinds.

It is originally the outer membrane, covering the lobes of the seed. Even there, as in the branch of a tree, it appears in the form of a kind of spunge, or like a crust of bread, composed of flatted bladders.

Its spunge-like nature may be further inferred from the contraction of its pores when dry, and the ease with which they dilate when in water. Grew has called it a most curious and exquisitely fine, wrought spunge. In the course of its growth, the outer ranges of these bladders drying, it becomes what we call the rind; for the rind was once bark, and has only suffered a slight change in separating from it.

By the bark the tree is fed with a continual supply of moisture, protected from external injuries, and defended from the excesses of heat and cold; for these purposes it is variously disposed in different trees. In the hardy and slow growing, as the oak and chesnut, it is thin; in the quick growing, as willow, poplar, and the like, it is thick. And what is more particularly to be attended to is, that in some its inner verge is radiated. There are some trees, and a great many herbaceous plants, in which this part is continued inward, in form of rays, through the blea into the wood, and seems to form so many green wedges, that split as it were the substance of both those parts;[135] a circumstance which accounts for the vegetation of some particular trees, which are known to live when deprived of the bark; because they have rays of the same substance within which answer the purpose, and this in a degree answering to the nature of their life.

The bark appears to be formed, first, of longitudinal fibres, which Du Hamel considers as so many lymphatic vessels; secondly, by a sort of a filmy cellular tissue, which has been considered as a kind of bladders by some, or as parenchymatous by others; thirdly, of the vasa propria interiora, or interior juice vessels.

The longitudinal fibres are disposed in strata, which lie one over the other. In that stratum which is next the rind, or rather the cellular coat, we perceive a net of longitudinal fibres, the meshes of which are large and easily distinguished, particularly when the cellular tissue that fills up the interstices is removed. To do this, the branches should be macerated for a considerable time; some require to be kept in this state for years. It will then be easy to separate first the rind, then the cellular coating, and afterwards this pulpy matter. It may sometimes be easily removed after the branches have been boiled.

The most exterior stratum, when examined by the naked eye, seems to be formed of simple fibres, which graft, solder, or inosculate one with the other; but when examined by a microscope, each of these fibres will be found to be a bundle of filaments, which may be easily separated from each other.

Grew says, that each filament, like the nerves in animals, consists of twenty or thirty small contiguous tubes, which run uniformly from the extremity of the root, without sending off any branches, or suffering any change in their size and shape. Hence the bark may be torn or divided lengthwise, with greater ease than in an horizontal direction; when macerated, they are capable of a very great degree of subdivision.

The filaments of a cortical vessel are to be looked on, agreeable to what we have already observed, as so many little bundles placed near together, and at first growing parallel to each other; but soon quitting this direction, the filaments of one fascicle parting from that to which they originally belonged, and inclining more or less obliquely towards another, sometimes uniting with it, at others, bending backwards, and uniting again with that from which it proceeded, or with some one that it meets with. In this manner new fascicles are often formed, while other parcels are increased or diminished by the additions of new filaments; by these means, a kind of irregular net is formed, and the fibres proceed in a serpentine line from the top to the bottom of the tree.

The thickness of the bark is entirely formed of strata of these longitudinal fibres, which lie one over the other; each of these strata is similar to the exterior one, only the meshes are smaller, and the fibres finer, in proportion as they are more interior, insomuch that at last the meshes are almost annihilated, and the fibres seem to lie quite parallel to each other.

There are some trees, however, where the meshes are not visible, and in which the fibres lie quite in a straight direction. There are many other circumstances in which they vary in different trees; in some the meshes of each stratum correspond with each other, diminishing gradually in size as they are more interior, and forming as it were so many conical cells.

We may, I think, conclude from what has been said, that the bark is composed of several thin membranes, which extend over the whole exterior surface of the tree. The most exterior membrane is the rind; under this is what Du Hamel calls the cellular coat; next to this the cortical stratum or true bark of the tree, which is formed of lymphatic vessels ranged more or less in a reticular form, and of the vasa propria interiora. The meshes are so constituted as to form large cavities next the rind, and small ones near the wood. These cavities are filled with a parenchymatous substance or the cellular tissue, which being continued from the wood to the rind, joins and unites the cortical stratum, and afterwards spreading on the outside thereof, forms what has been termed the cellular coat.

OF THE CELLULAR TISSUE.

We now proceed to give some account of the substance which fills up the vacant spaces that are left between the longitudinal fibres. It is called by Grew the parenchyma or pulp, by Malpighi, the vesicular tissue or web; both of them consider it as formed of small bladders or reticles, that are in contact with each other, lying in an horizontal position, or at right angles to the longitudinal fibres: they do not suppose them to be all of the same size, or even of the same figure: Grew compares it to the froth of beer or eggs. The flesh of fruits consists for the most part of this substance, very much filled with juice, though with considerable difference in its organization. Be this as it may, the nature of this substance, its form and structure, are at present but very little known. It is floccose, and varies in colour in different species.

OF THE VASA PROPRIA INTERIORA.

Besides the lymphatic vessels and the cellular substance, we find the juice vessels, or vasa propria, in the bark. In those trees which are famous for medicinal virtues, they are usually very large; they carry the milky juices of the sumach, and in them is lodged the finest and highest-flavoured turpentine in all the kinds of pine. Dr. Hill thinks that a tree of that genus exhibits them best, and the more, as the turpentine which fills them may be perfectly dissolved in spirit of wine. The pinus orientalis is the species in which these vessels are most distinctly seen.

OF THE BLEA.

This is that part of the tree which is formed into wood, and therefore lies between it and the bark, and may be separated from them by maceration.

A longitudinal piece of the blea, when examined by the microscope, exhibits a number of vessels running parallel to each other, the interstitial spaces being filled with a floccose, white, formless substance, of which Dr. Hill suspects even the vessels themselves to be formed. Innumerable small openings or mouths may be discovered in these vessels, suited to imbibe the moisture which is so essential to the life and health of plants. These mouths cannot be well discerned, except when they are opened by the season of the year, either before the first leaves of spring, or in the midsummer shooting time; though a small quantity of moisture will keep them open at that time, yet no quantity would be sufficient at an improper season.[136]

The blea is a zone more or less perfect, which lies under the bark, and covers or surrounds the wood, and is principally distinguished from it by being less dense. In some species the difference between the blea and the wood is very remarkable, in others it is less so.

The ancient botanists, struck with the difference they observed between the wood and the blea, compared this substance to the fat in animals. Malpighi, Grew, and Du Hamel considered it as the wood not yet arrived to a state of perfection. It is organized in a manner similar to the wood, and possessing the same vessels disposed nearly in the same manner. The juice vessels of this part may be separated from it by maceration; Dr. Hill says, that in this state they appear perfect cylinders, with thick white coats, the surface perfectly uniform.

OF THE WOOD.

When the bark and the blea are taken away, we come to the wood, which is a solid substance, on which the strength of the tree depends, and which has been considered by naturalists as being to the tree what bones are to the animal. The wood, in a general view may be considered as formed of strata, which are inclosed one within the other; these strata consist of ligneous fibres or lymphatic vessels, the cellular web or tissue, vasa propria, and what have been called the air vessels. It is more difficult to investigate the construction of the wood than that of the other parts, because the texture is in general much harder, and therefore not so easily separated, requiring very long macerations, and many subjects, before one may be found fit for examination.

If a transverse section of almost any kind of wood be examined, we shall perceive these strata very clearly and sensibly distinguished from one another. It has been generally supposed that each of these is the product of one year’s growth; though, if we cut the same wood obliquely, it will be found that each of these strata is compounded of smaller ones, which are therefore not so easy to discover as the larger. By macerating rotten pieces of trees, the wood may be divided into an immense number of leaves or strata, thinner than the finest paper.

If the foregoing strata be examined in their detached state by the microscope, we shall find them to be composed of longitudinal fibres; some pieces of rotten wood, after maceration, will divide of themselves into very fine longitudinal fibres; the existence of these is further proved by the facility with which wood may be split in the direction of these fibres. From hence we may collect, that the ligneous strata are formed of small fibres or vessels, collected together in fascicles, like the bark: in some trees they are parallel to each other, in others they are disposed more obliquely, crossing and forming an irregular kind of network. There is great probability that this reticular disposition exists in all trees, though it may be difficult to discover it in many on account of the fineness of the meshes, the hardness of the wood, and the sameness of colour in the constituent fibres.

We are here only speaking of the lymphatic vessels or ligneous fibres of the wood, which exist in it as well as in the bark, though in different states; for the ligneous fibres are always harder and less flexible than the cortical ones. Malpighi thinks they differ in another particular, namely, that a juice or fluid issues from the cortical fibres, while none is found in those of the wood. In this it would appear from the observations of Du Hamel, that he was mistaken.

A transverse section of wood generally appears formed of a number of rays proceeding from the corona to the bark, which are intersected at different distances by concentric circles, interspersed with vessels of varying magnitude: the variations in this structure afford much pleasure to the curious observer, and throw considerable light upon the nature and properties of timber; for it is by means of a variety of strainers that different juices are prepared from the same mass. Matter, considered as matter, has no share in the qualities of bodies. It is from the arrangement of it, or the recipient forms given to it, that we have so many different substances. According to the modifications that these receive, we shall find the same light, air, water, and earth, manifesting themselves in one by a deadly poison, and in another by the most salubrious food. A lemon ingrafted upon an orange stock, is capable of changing the sap of the orange into its own nature, by a different arrangement of the nutritive juices. One mass of earth will give life and vigour to the bitter aloe, to the sweet cane, the cool house-leek, and the fiery mustard, the nourishing grain, and the deadly night-shade.

The wood may be considered as composed of two parts, ligneous and parenchymatous. The former has already been treated of; the latter is that which is disposed into rays, running as it were between the ligneous fibres, and interweaving with them; it originates either with the pith or corona. There is a very great diversity in these radial insertions; in some trees there are very few, while they abound in others; in some they are very fine, in others very thick. In texture, they seem similar to the blebs of the bark, only that here they are so crowded and stretched out as to appear like parallel threads, somewhat similar to a net when drawn tight.

OF THE CORONA.

Dr. Hill gives this name to that circle which surrounds the pith, and separates it from the wood; although in his opinion it differs greatly from both, and in its composition has no resemblance to either. It is, according to him, the most important part in the whole vegetable fabric, by which the propagation and increase of the branches, buds, and shoots, are carried on.[137]

It has been usual to suppose the pith of vegetables to be the part in which these wonderful sources of increase reside, but this is not the case; and he asserts, that so far from being prior to the other parts, it is in reality posterior to some of them in the time of its formation.

The corona is not so uniform as the other parts, nor is it constituted exactly similar in all trees. It is placed between the pith and wood in all vegetables, forming a ring, whose outline is more or less regulated. The general circle is cellular, composed of blebs and vessels, like the bark and the rind, and is perfectly similar to them, only that at different distances oblong clusters of different vessels are placed amongst it. These clusters are usually eight or ten in number, and give origin to the angles of the corona. They are not uniform, or of one kind of vessels, as in the bark, but each has two distinct sorts, the exterior one answering to the blea, and the interior, to the wood of trees; and within each of these are disposed vessels not unlike those in the blea and wood, though often larger than they are found in those parts.

Thus each cluster is composed of all the essential parts of the succeeding branch, and the intermediate parts of the circle are absolutely bark and rind; they are ready to follow and clothe the cluster when it goes off in the form of a shoot, because it will then need their covering and defence, though in its present inclosed state it does not. It is from this construction, that a tree is ready at all times and in all parts to shoot out branches, and every branch in the same manner to send out others; for the whole trunk, and the branch in all its length, have this course of eight or ten clusters of essential vessels ready to be protruded out, and the proper and natural integuments as ready to cover them. In some trees, these parts are more evident, in others more obscurely arranged. Dr. Hill says, the bocconia, or parrot-wood of the West-Indies, and the greater celandine, are proper subjects for opening this great mystery of nature. On the corona and its clusters depend that property of vegetables, that they can be produced entire from every piece. These clusters follow the course of the other portions of the tree; they are therefore everywhere; they are always capable of growing, and their growth, even in a cutting of the smallest twig, cannot produce a leaf, or any other part of a vegetable alone, but must afford the whole; for they are complete bodies, and the whole is there waiting only for the opportunity of extension, by obtaining sufficient nourishment. For the knowledge we have of this part we are altogether indebted to Dr. Hill. It remains for future observers to confirm, or disprove his observations.

OF THE PITH.

The pith is found in the center of every young shoot of a tree; it is large in some, less in others, but present in all. It is placed close within the corona.

It seems to be nothing more than a congeries of the cellular tissue; it is generally found near the center of the tree, inclosed as it were within a tube; in general, the cells of the pith are larger than those of the cellular tissue, with which, according to Du Hamel, it communicates. For the rays which extend from the pith to the bark are, in his opinion, produced from it. Thus, though it may differ in name from the parenchymatous parts of the bark, and the radial insertions in the wood, yet it is of the same nature and texture, and is continuous with them; so that, according to this idea, the skin, the parenchyma, the insertions, and the pith, are all one piece of work, filled up in divers manners with the vessels.

The bark and the wood grow thicker every year, while the pith, on the contrary, grows more slender, so that in a branch of one year it is of a larger size than it is in the same branch when two years old, and so on. In very young branches, while in an herbaceous state, the pith forms the greatest part of its substance; but when the fibres are stronger, the pith becomes less succulent, and surrounded with a tube of wood; when the branch has arrived to a certain age, it is so compressed as to be almost annihilated. In examining different branches that proceed from others in their first state, a small communication between the pith of the one and the other will be found; but this communication is generally entirely closed up in the second or third year.[138] The cells of which the pith is formed are at first entirely one connected body; but as the plant grows up, it is often so broken and ruptured, as to remain no longer a continuous substance.

This, as well as many other particulars in the history of the pith, corroborates the opinion of Dr. Hill,[139] who thinks it is formed for the purpose of moistening the clusters of the corona, and regulating its extension; it has been supposed coeval with, or primordial to all the other parts, but he thinks it is postnate, and comes after them in the order of time, as well as in its uses; that exhaled air gives origin to its blebs, and the thickness of the juices cloathing the bubble, gives it form and substance. The first season is the time of its greatest use, and it immediately after begins to decay.

The pith has in general been represented as much more complex than it really is. It consists of a range of bladders lying one over the other. The membrane is simple, the outline single; but as it is very difficult to procure it in this simple state, it is often seen and represented under a variety of irregular, though pleasing forms, which are occasioned by the intersections of the outlines of the blebs, as seen one over another.

A cluster in any part of the corona, protruding itself onward and outward in the growing season,[140] carries a part of the circle out with it. The cluster itself is a perfect piece of the wood and blea, and the bark which follows it out in its progress perfectly clothes it; thus is the first protrusion of the shoot made, but all this while there is no pith. The continuation of growth is made by the extension of all the parts obliquely upwards; in the course of this extension they hollow themselves into a kind of cylinder, of the form of the future branch, and by this disposition a small vacancy is made in their center. This enlarges as they increase, and as it enlarges it becomes filled with the exudation of those little bladders which remain and constitute the pith, fed from the inner coat of the pith, which already begins to form itself into a new corona. Grew seemed to think, that in some instances the pith was of posterior growth to the other parts, and derived its origin from the bark; and that the insertions of the bark running in between the rays of the wood meet in the center, and constitute the pith.

OF THE SAP VESSELS.

The most numerous and the largest apertures are generally to be found in the wood, which are perceived very distinctly in a transverse section, in which the ends of the vessels are seen as cut through by the knife. The scarlet oak of America is recommended as a proper object for exhibiting them. If a short cylinder of a three years branch of this oak, a little macerated, be hollowed away with a chissel, we shall see what a large portion of the wood is occupied by these vessels; they are thick and strong, and it is easy, with some care and attention, to loosen several of them.

If a number of these thus separated be put into a vial of rain water, and frequently shook for several days, some will at length be found perfectly clean; these are then to be put into spirit of wine, and when that has been two or three times changed, they will be in a condition to be viewed for understanding their structure; another method of preparation has already been shewn in page 162.

These are the vessels which have been called by some writers air, by others, tracheal vessels. It is, however, to be remarked, that most of those who have considered them as air vessels, refer us to the tree while in a more herbaceous state; in this case they say, that we shall find these parts filled with a fine spiral filament. As these vessels are often to be found empty, they have been supposed to answer the purposes of lungs to the plant. Malpighi asserts, that if they be examined in winter, they often exhibit a vermicular motion, which astonishes the spectator.

Those who suppose the corona to contain the whole structure of the tree in miniature, and that it is the embryo of future shoots, suppose it to contain the vessels proper for each part, a subject that must be left to the decision of future observers.

OF THE VASA PROPRIA INTIMA.

These are the only vessels which remain to be spoken of. They are large, conspicuous, and important; their natural place is in the blea, though they are sometimes repeated in the wood and the corona. Their coats are thicker than those of any other vessels.[141] It is not difficult, after a successful maceration, to separate some of these vessels from the blea; in this state they appear perfect cylinders, with thick white coats, of a firm, solid, and uniform texture.

It has generally been supposed, that each of those concentric circles, which are to be observed in the transverse section of almost every tree, was the product of one year, or the quantity of wood added to the tree in that space; here, however, Dr. Hill differs again from the general opinion.

From what has been said, we may deduce the following general ideas relative to the organization of trees. The most obvious and remarkable parts of a plant, or tree, are the root, the stem, the branches, the leaves, the flower, and the fruit. The component parts of these divisions are not complicated; they are simple when compared with those of an animal, and this because the offices of the vegetable are fewer than those of the animal.

The interior part may be considered as consisting of ligneous fibres, interspersed with a vast number of bladders, which are here named the cellular tissue, the vasa propria, and the sap vessels; though these are considered by some writers as mere air vessels.

The ligneous fibres are very fine tubes, proceeding nearly in a vertical direction from the top to the bottom of the tree; they are sometimes parallel to each other, sometimes they divaricate, and often leave oblong intervals or spaces. There is great reason for supposing them to be a species of lymphatic vessels. The vacant spaces between these fibres are filled up by a vesicular membrane, lying in an horizontal direction, and which is called in this chapter the cellular tissue.

The vasa propria are formed of ligneous fibres, but differ from the foregoing in their size, and in the juices which they contain. In the part properly called the wood, we meet with the sap vessels; but as in some states they seem as if they were formed of a silver-coloured spiral membrane, and are found without any juices, they have been supposed to be air vessels, and called the trachea, making up an arterial system, and supplying the place of the heart in animals.

The interior part of the tree may be further considered as divided into four principal concentric strata, the bark, the blea, the wood, and the pith; to these Dr. Hill has added the corona. Whatever part of a plant is examined, we find these and no more. The root, its ascending stalk, and descending fibre, are formed of one, and not three different substances. Thus the whole vegetable is reduced to one entire body. And what appears in the flower to be formed of altogether distinct parts, will be found to originate in these.

The bark, which is the exterior covering of the tree, is divided into two parts, a thin outer rind, and a much thicker inner one. The exterior one seems to be little more than a fine film of irregular meshes, the inner one composed of large blebs, leaving in some subjects large vacant spaces, which form its vasa propria. It is made up of several strata lying one over the other.

Next to this is the blea, which is of an uniform structure. It is an imperfect wood, waiting only for the hand of time to be brought to perfection. The duration of the blea in this middle state depends on the internal powers and strength of the tree, being so much shorter as this is more vigorous.

The wood, including the corona, comes next; it differs in density and duration both from the blea, the bark, and the wood. It is made up of strong fibres. The life of the vegetable seems to reside in it; from it all the other parts are produced. It shoots a pith inwards, and a blea and a bark outwards.

Every tree may be considered as consisting of numerous concentric strata or flakes, forming so many cones, inscribed one within the other, and whose number is almost indefinite. The most exterior contain the rudiments of the bark; the more interior, those of the wood. In the germ they are gelatinous, by degrees they become herbaceous, and in process of time assume the consistence of wood. Thus the stem, the root and the branch, may be considered as formed of a prodigious number of concentric vertical strata, each composed of different fascicles of fibres; which fibres are again formed of smaller ones. The spaces between these, and among the fibres, are filled up, interwoven with, and connected by the cellular tissue, of which the radial insertions are formed.

The strata harden successively one after the other; the most interior stratum is that which hardens first; this is then covered by another which is more ductile and herbaceous, and so on; so that the bulk of the tree is increased every year by the accession of an hollow cylinder of wood derived from the internal bark. From the extension in breadth, the tree acquires bulk; from that in length it gains its height. The strata gradually diminish in size as they gain in length; from hence the conical figure of the root, stem, and branch. All the parts of the plant are the same, differing in nothing more than in shape and size. The roots are sharp and pointed, that they may make their way more readily through the earth. The leaves are broad, that they may more effectually catch the moisture from the atmosphere, &c. When the root of a tree is elevated above, instead of being retained under the earth, it assumes the appearance of a perfect plant, with leaves and branches. Experiment shews that a young tree may have its branches placed in the earth, and its roots elevated in the air, and in that inverted state it will continue to live and grow. The principal source of the phÆnomena of vegetation is the simplicity and uniformity of their organization.

The figures in Plates XXVIII. XXIX. and XXX. are portions of transverse sections of trees and herbs. The sections were cut by Mr. Custance,[142] who first brought this art to perfection, and remains hitherto unrivalled in these performances.

Plate XXVIII. Fig. 1, exhibits a piece of an herb growing on rubbish, and known by the name of fat-hen:[143] Fig. 2, a microscopic view of the same. Fig. 3, a magnified representation of a section of a reed that comes from Portugal: Fig. 4, the real size of the section.

Plate XXIX. Fig. 1, is a magnified view of a section of the althea frutex: Fig. 2, the natural size of the section. Fig. 3, a magnified view of a section of the hazel: Fig. 4, its natural size. Fig. 5, a microscopic view of a section of a branch of the lime-tree: Fig. 6 represents its natural size.

Plate XXX. Fig. 1, a magnified view of a section of the sugarcane: Fig. 2, its natural size. Fig. 3, a magnified view of a section of the bamboo cane: Fig. 4, the natural size. Fig. 5, a magnified view of a section of the common cane: Fig. 6, the real size.